Cockerell 1905 : 131 Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos Choi, Jinyeong Soysouvanh, Pheophanh Lee, Seunghwan Hong, Ki-Jeong Zootaxa 2018 2018-08-17 4460 1 1 62 Cockerell, 1905 COckerell 1905 [151,621,965,991] Magnoliopsida Apocynaceae Pulvinaria GBIF Plantae Gentianales 45 46 Tracheophyta species polygonata     Pulvinaria polygonata  Cockerell 1905: 131.   Diagnosis.Dorsal derm with polygonal reticulations; tubular ducts present, but small ( Fig. 42E); duct tubercles present ( Fig. 42D). Marginal setae bluntly spinose, mostly with fimbriate apices ( Fig. 42A). Stigmatic clefts distinct, each containing 4 or 5 stigmatic spines ( Figs 41C, 42C). Venter with multilocular disc-pores usually each with 8 loculi, abundant around vulvar area, a few pores also present laterad of meta- and mesocoxa ( Fig. 42I); tubular ducts of 3 types: type I each with a broad inner ductule, present on medial area of head, thorax and anterior abdomen; type II each with a narrow inner ductule, present on medial and inner submarginal area of head, thorax and abdomen; and type III each with a filamentous inner ductule, present in submarginal areas ( Figs 41D, 42J); antenna 8 segmented (partially adopted from Williams & Watson, 1990).    Material examined. 10 ♀♀, LAOS, Kham Dist., Xiangkhoang Prov.,  2.v.2015, coll. J.Y. Choi, on  Citrussp. ( Rutaceae).   Hosts.Polyphagous. According to García Morales et al. (2016),  P. polygonatahas been recorded from plants belonging to 10 genera in 7 families. In Laos, it has been found on  Ficussp. ( Moraceae) ( Suh & Bombay2015).   Distribution.Mainly known from Australian, Oriental( Bangladesh, India, Laos, Philippines,  SriLanka, Taiwanand Vietnam) and Palearctic Regions ( Suh & Bombay2015; García Morales et al. 2016).  Economic importance. Mani & Krishnamoorthy (1998)considered  P. polygonatato be a serious pest of mango (  Mangifera indica) in India. In addition, Williams & Watson (1990)noted that  P. polygonatacould be a potential pest of citrus because of its host preferences. The species is frequently intercepted at U.S. ports ( Miller & Miller 2003).   Remarks.  Pulvinaria polygonataclosely resembles  P. aurantiiCockerell, but is easily separated by the number of stigmatic spines in each spiracular cleft:  P. polygonatahas 4–5, whereas  P. aurantiihas only 3 ( Takahashi 1955b). 1918155552 2015-05-02 J. Y. Choi & Citrus Laos Kham Dist. Xiangkhoang Prov. 46 47 10 10