Schiner, 1868 : 255 Enderlein 1922 : 228 Hennig 1935 :49 Plocoscelus niger ( Schiner, 1868 ) Hennig 1935 : 49 A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae) Ferro, Gustavo Borges Marshall, Stephen A. Zootaxa 2018 2018-06-07 4429 3 401 458 8BBX5 Schiner, 1868 Schiner 1868 [151,575,151,178] Insecta Micropezidae Cardiacephala GBIF Animalia Diptera 34 435 Arthropoda species nigra     Cardiacephala nigra  Schiner, 1868: 255.    Rhoecius vulgus  Enderlein 1922: 228,  Hennig 1935:49.    Plocoscelus niger( Schiner, 1868),  Hennig 1935: 49.   Description:Body length 9 mm, wing 6 mm. Palpus mostly black, white on outer face. Clypeus black and shiny with some sparse pale microsetulae. First flagellomere black with pale microsetulae. Fronto-orbital plates and frontal vitta dark brown. Hind femur narrowly white basally. Row of katepisternal setae brownish. Fore tarsus dark brown with gold/whitish pubescence ventrally at base of tarsomere one. Oviscape black, shiny. Epandrium brownish. Head: Palpus broadened basally and gradually tapering to apex. Clypeus short, length less than half of width. Two frontal setae. Frontal vitta with a narrowed depression centrally, reaching ocellar triangle. Epicephalon slightly depressed. Inner vertical seta insertion flat, without tubercle. Paracephalon smooth, without swelling (not extending beyond the vertex). Vertex flattened. Line of black supracervical setae strong. Thorax: Mesonotum with white pruinosity. Scutellum upturned, triangular. Femora with sparse yellowish setulae. Mid and hind tibiae flattened, sulcus on outer face very pronounced and curved ( Fig. 55). Wing with stigmatal and discal bands, apex entirely dark. Crossvein r-m clear. R4+5 and M1+2 meeting at the wing margin. A1+CuA2 as long as CuA2. Female cervical sclerite smooth, without swelling. Abdomen: Tergites covered by black setulae, some of them shiny gold. Female abdomen: Single spermathecal duct apically broad, spermatheca cylindrical and apically invaginated. Paired spermathecal duct delimited by a narrow basal third and broader distal two thirds. Paired spermathecal duct stems smooth and thin in basal third, followed by a small rounded swelling and a broad apical third ( Fig. 56). Paired spermathecae striate and sac-like, distally swollen and tapered basally. Male abdomen: Ejaculatory apodeme bigger than epandrium and with a broad blade, wider than sperm pump ( Fig. 58). Phallapodeme extending to anterior limit of hypandrium. Distiphallus longer than epandrium, ending in a phallic bulb, phallic bulb almost spherical and somewhat broader than the basal distiphallus.    Typematerial: Holotypeof  Cardiacephala nigra( 1 ♂, NMW): “South America”. Date and collector not cited; Syntypesof  Rhoecius vulgusEnderlein( 2 ♂, ZMHB): BOLIVIA. Santa Cruz, Prov. Sara. J. Steinbach. COLOMBIA. Moritz coll.  Material examined: BOLIVIA. 1 kmE Guanay, 580 m, 15º30.2’ S 69º52.3’ W, 13.III.2001, S. D. Gaimari, ex: human excrement ( 2 ♀, USNM); 1 kmE Guanay, 580 m, 15º30.2’ S 69º52.3’ W, 14.III.2001, S. D. Gaimari ( 1 ♀, USNM); La Paz, Guanay, 40 kmS roadside, 12.IV.2001, S. A. Marshall ( 1 ♀, DEBU00149543); La Paz, Mapiri 5 kmW, 750 m, 15º17.8’ S 68º15.6’ W, 16.III.2001, Amnon Freidberg ( 1 ♀, USNM); La Paz, Mapiri, bank of Rio Mapiri, 15º18’34” S 68º13’ W, 12.IV.2001, A. L. Norrbom & G. A. Kung ( 1 ♂, USNM); La Paz. Rio Merke (= Rio Chiñijo), 3 kmNW Mapiri, 15º18’48” S 68º14’25” W, disturbed sec. forest, 9.IV.2001, S. A. Marshall ( 1 ♂, DEBU00150492); La Paz, San Juanito nr. Teoponte, 500 m, 15º29’42S 67º47’48W, 8.IV.2001, S. A. Marshall ( 5 ♀, 1 ♂, DEBU00149601, DEBU00149602, DEBU00149632, DEBU00149645, DEBU00149647, DEBU00150734); La Paz, San Juanito nr. Teoponte, 500 m, 15º29’42S 67º47’48W, 8.IV.2001, S. A. Marshall ( 1 ♀; DEBU00149662, sequence MYCRO240–16); La Paz, Tajihui, 590 m, 15º40.8’ S 67º41.7’ W, 12.III.2001, Amnon Freidberg ( 2 ♀, 1 ♂, USNM); BRAZIL. Amazonas, Manaus, R. Ducke, Arm. Malaise, 21.VII.1990, J. Vidal & R. A. Rocha ( 1 ♀, INPA); Pará, Tucuruí, 7.VIII.1980, R. S. Mansão ( 1 ♀, CNC); São Paulo, Sorocaba, 10.IX.2011, G. Ferro ( 1 ♀, DZUP); COLOMBIA. Magdalena, Tayrona Natl. Pk. Zaino, 50 m, malaise trap, 28.IV–13.V.2000, R. Henriquez ( 3 ♀, 2 ♂, DEBU00138497, DEBU00138501, DEBU00138503, DEBU00138507; DEBU00138508, IAVH); Magdalena, Tayrona Natl. Pk., Admin. Cañaveral, Malaise trap, 30 m, Malaise, 26.IV–12.V.2000, R. Henriquez ( 1 ♀, 1 ♂, DEBU00138437, DEBU00138438, IAVH); Magdalena, PNN Tayrona Cañaveral, 11º20’ N 74º2’ W, 30 m, Malaise 11/03/00 1/2/01R, M. 963, R. Henriquez ( 1 ♀, 1 ♂, IAVH); Magdalena, Tayrona Zaino, 50 m, 11º20’1” N 74º2’1” W, 29.V–14.VI.2000, M. 241, R. Henriquez ( 2 ♂, IAVH); Magdalena, Tayrona Zaino, 50 m, 11º20’1” N 74º2’1” W, 28.V–17.VII.2000, M. 301, R. Henriquez ( 1 ♀, IAVH); Magdalena, Tayrona Zaino, 50 m, 11º20’1” N 74º2’1” W, 29.IX–17.X.2000, M. 793, R. Henriquez ( 1 ♀, IAVH); Magdalena, PNN Tayrona Cañaveral, 11º20’ N 74º2’ W, 30 m, Malaise 11/03/00 11/22/00R, M. 943, R. Henriquez ( 2 ♀, IAVH); Magdalena, Tayrona Zaino, 50 m, Malaise trap, 11º20’1” N 74º2’1” W, 28.IV–13.VI.2000, R. Henriquez. ( 1 ♂, DEBU00138496, IAVH, sequence MYCRO241–16); Putumayo, PNN La Paya Cabaña Viviano Varzea, 0º7’ S 74º56’ W, malaise, 320 m, 15– 31.X.2001, M. 2438, R. Cobete ( 1 ♀, IAVH); COSTA RICA. Estrella Valley, Pandora, 16.III.1984, FIT, GV Manley & HF Howden ( 1 ♀, DEBU); Heredia, Estac. Biol. La Selva, 15.II.1994, mal. trap, M/02/349, ALAS( 1 ♀, MNCR); Heredia, Estac. Biol. La Selva, 1.III.1994, mal. trap, M/01/360, ALAS( 1 ♀, MNCR); Heredia, La Selva Biol. Res. Nr. Puerto Viejo, mainly trail, 9–10.II.2003, M. Pollet ( 1 ♀, 1 ♂, DEBU00241584, DEBU00241593, IRSNB); Heredia, La Selva Biol. Stn., 10º26’ N 84º01’ W, 150 m, Trail 0-400 STR., 9.II.2003, M. Pollet ( 1 ♀, DEBU00241580, IRSNB); Prov. Punt. Est. Queb. Bonita, 50 m, reserv. Biol. Carara, II.1991, L-N-194500 469850, C. Moraga ( 1 ♂, DEBU); San Jose, Pérez, Zeledón, Rivas, 820 m, 27.XII.1998– 5.I.1999, B. D. Gill ( 1 ♂, DEBU00151033); ECUADOR. Limoncocha, 250 m, 0º24’ S 76º40’ W, 9–16.III.1976, G. E. Shewell ( 1 ♀, CNC503365); Napo, Coca, NapoR., 250 m, V.1965, L. Pena ( 63 ♀, 2 ♂, CNC503338, CNC503339, CNC503340, CNC503341, CNC503351, CNC503352, CNC503353, CNC503354; CNC503356, CNC503357, CNC503359, CNC503360, CNC503361, CNC503362, CNC503363, CNC503367, CNC503369, CNC503371, CNC503373, CNC503374, CNC503375, CNC503377, CNC503378, CNC503379, CNC503380, CNC503382, CNC503383, CNC503384, CNC503385, CNC503386, CNC503387, CNC503388, CNC503389, CNC503390, CNC503391, CNC503392, CNC503393, CNC503394, CNC503395, CNC503396, CNC503397, CNC503398, CNC503399, CNC503402, CNC503403, CNC503404, CNC503405, CNC503406, CNC503407, CNC503408, CNC503409, CNC503410, CNC503411, CNC503412, CNC503414, CNC503421, CNC503422, CNC503423, CNC503425, CNC503426, CNC503428, CNC503430, CNC503434, CNC503436, CNC503438); Napo. Coca. NapoR.,  250 m. 25–30.IV.1965. L. Pena ( 1 ♀, 1 ♂, CNC503344, CNC503366); Napo, Jatun Sacha Biol. Res. 6 kmE Misahualli, 1º4’ S 77º37’ W, 450 m, 30.IV–8.V.2002, S. A. Marshall ( 13 ♀, 2 ♂, DEBU00178784, DEBU00178805, DEBU00178806, DEBU00178816, DEBU00178822, DEBU00178947, DEBU00178991, DEBU00178998, DEBU00179007, DEBU00179008, DEBU00179009, DEBU00179016, DEBU00179062, DEBU00179130, DEBU00179160); Napo, Jatun Sacha Biol. Res. 6 kmE Misahualli, 1º4’ S 77º37’ W, 450 m, 29.IV–8.V.2002, K. Peiman ( 1 ♀, DEBU); Napo, Jatun Sacha Biol, Res. 6 kmE Misahualli, 1º4’ S 77º37’ W, 450 m, 1–2.V.2002, M. Buck ( 1 ♀, 1 ♂, DEBU00178207, DEBU00178208); Napo, Jatun Sacha Biol. Res. 6 kmE Misahualli, 1º4’ S 77º37’ W, 450 m, 1–2.V.2002, M. Buck & Lonsdale ( 2 ♀, DEBU00186979, DEBU00186980); Napo. Jatun Sacha Biol. Res. 6 kmE Misahualli. 1º4’ S, 77º37’ W,  450 m. 2.V.2002. M. Buck ( 3 ♀, DEBU00177943, DEBU00177947, DEBU00177948); Napo, Jatun Sacha Biol. Res. 6 kmE Misahualli, 1º4’ S 77º37’ W, 450 m, 2– 7.V.2002, M. Buck ( 1 ♀, DEBU00187281); Napo, Jatun Sacha Biol. Res. 6 kmE Misahualli, 1º4’ S 77º37’ W, 450 m, 30.IV–7.V.2002, Landslide in forest, M. Buck et al.( 3 ♀, DEBU00196363, DEBU00196364, DEBU00196367); Napo, Jatun Sacha Biol. Res. 6 kmE Misahualli, 1º4’ S 77º37’ W, 450 m, 7.V.2002, M. Buck ( 2 ♀, DEBU00178866, DEBU00178868); Napo, vic. Misahualli ca., 1º2’ S 77º40’ W, 450 m, 27–31.X.2002, E. M. Fisher ( 4 ♀, DEBU); Napo, Jatun Sacha, 7.V.2002, S. A. Marshall ( 1 ♀, DEBU); Napo, Jatun Sacha, 5.V.2002, S. A. Marshall ( 1 ♂, DEBU); Pompeya, NapoR. Pastaza, 250 m, 14–22.IV.1965, L. Pena ( 2 ♀, CNC503349, CNC503424); Rio Palenque, (Univ. Miami Res. Sta.), 29.VII.1978, G. J. Umphrey ( 1 ♂, DEBU); HONDURAS. Gracas a Dios, Rio Paluca, Kausirpe, 22–24.V.1994, B. D. Gill ( 1 ♀, DEBU00138357);  MEXICO. Pue. Iz. Metamoros, 26.VIII.1969, L. A. Kelton ( 1 ♀, CNC503343); Tabasco, Teapa ( 8 kmSW), 6.V.1985, Amnon Freidberg ( 4 ♀, 3 ♂, TAU);  PERU. Avispas, Madre de Dios, 400 m, 10–20.IX.1962, L. Pena ( 3 ♂, CNC503345, CNC503346, CNC503364); Cusco, Estacion Biologica Villa Carmen, 12º53’39”S 71º24’14”W, 540m, 12– 14.XII.2013, A. L. Norrbom & B. D. Sutton ( 2 ♂, USNM); Madre de Dios, Pantiacolla Lodge, Alto Madre de DiosRiver, 420 m, FIT, 12º39’18” S 71º13’54” W, 14–19.XI.2007, D. Brzoska ( 2 ♀, DEBU00343608, DEBU00356554); TRINIDAD. Tabaquite, 27.XI.1902, H. D. Chipman ( 1 ♀, CMNH); Tabaquite, 2.XII.1902, H. D. Chipman ( 1 ♀, CMNH); Tabaquite, 19.XII.1902, H. D. Chipman ( 1 ♀, CMNH); Tabaquite, 22.XII.1902, H. D. Chipman ( 1 ♀, CMNH); Tabaquite, 7.I.1903, H. D. Chipman ( 3 ♀, CMNH); Tabaquite, 20.I.1903, H. D. Chipman ( 2 ♀, CMNH); Tabaquite, 10.XI.1903, H. D. Chipman ( 2 ♀, CMNH); Tabaquite, 12.XI.1903, H. D. Chipman ( 1 ♀, CMNH); Tabaquite, 21.XI.1903, H. D. Chipman ( 1 ♀, CMNH); Tabaquite, 20.XI.1903, H. D. Chipman ( 1 ♀, CMNH); VENEZUELA. Sucre, Puy-Puy, 32kmE Carupano, plantations & 2º forest nr. beach, sweep, 31.III– 2.IV.1988, M. von Tschirnhaus ( 1 ♂, DEBU00256927)   Distribution: Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Honduras,  Mexico, Peru, Trinidad, Venezuela.