Schiner 1868 : 236 Heterochroa picta Schiner 1868 philippii Rondani 1863 Heterochroa Boisduval (Nymphalidae) Marschall 1873 : 333 Scudder 1884 : 147 Schulze et al . 1930 : 1534 Czerny 1903 : 98 Peratochaetus Czerny 1903 : 98 Heterochroa bicolor Schiner 1868 Soós 1968 : 2 Kertész 1906 : 321 Melander & Argo 1924 : 6 Schulze et al . 1926b : 233 Malloch 1933 : 238 Neave 1939a : 251 Stuardo Ortiz 1946 : 150 Hennig 1952 : 22 Clusiidae Frey 1960 : 28 McAlpine 1960 : 67 Soós 1964a : 5 Cole & Schlinger 1969 : 426 Lonsdale & Marshall 2008c : 8 Alloclusia Hendel Diptera Clusiinae Peratochaetus Peratochetus Rondani Clusia Haliday Philippii Rondani pictipennis Wulp ornata Johnson Heleomyzidae Peratochetus Rondani flava Meigen Clusia Czerny 1903 : 92 Heterochroa Schiner Peratochaetus Aldrich 1905 : 571 Spilochroa Williston Heleomyzidae Kertész 1906 : 321 Malloch 1933 : 234 Alloclusia Hendel 1917 : 36 Heterochroa Schiner Peratochetvs philippii Rondani 1863 Melander & Argo 1924 : 32 Schulze et al . 1926a : 108 Malloch 1933 : 238 Neave 1939a : 116 Stuardo Ortiz 1946 : 149 Hennig 1952 : 227 Clusiidae Frey 1960 : 28 McAlpine 1960 : 67 Soós 1964a : 5 Cole & Schlinger 1969 : 426 Lonsdale & Marshall 2008c : 8 Apiochaeta Clusiidae Hendelia beckeri Czerny Apiochaeta philippii (Rondani) Apiochaeta bicolor (Schiner) aterrima (Malloch) Malloch 1933 : 244 Frey 1960 : 28 Soós 1968 : 2 Lonsdale & Marshall 2008c : 9 aequalis Malloch Malloch 1933 : 245 Stuardo Ortiz 1946 : 150 Frey 1960 : 28 Soós 1968 : 2 Lonsdale & Marshall 2008c : 9 aterrima Malloch bicolor (Schiner) Schiner 1868 : 237 Handlirsch 1884 : 142 Clusia Haliday Heterochroa Schiner Apiochaeta bicolor Czerny 1903 : 98 Melander & Argo 1924 : 6 Malloch 1933 : 243 Stuardo Ortiz 1946 : 150 Frey 1960 : 28 Soós 1968 : 2 Lonsdale & Marshall 2008c : 11 chiloensis (Malloch) Malloch 1933 : 238 Malloch 1933 : 238 Stuardo Ortiz 1946 : 149 Frey 1960 : 28 Soós 1968 : 1 Apiochaeta chiloensis Lonsdale & Marshall 2008c : 12 Malloch 1933 : 244 Frey 1960 : 28 Soós 1968 : 2 Lonsdale & Marshall 2008c : 13 costalis (Malloch) Malloch 1933 : 239 Malloch 1933 : 239 Frey 1960 : 28 Soós 1968 : 2 Apiochaeta costalis Lonsdale & Marshall 2008c : 15 dacryodes Lonsdale & Marshall. Lonsdale & Marshall 2008c : 15 Lonsdale & Marshall 2008c : 15 excepta (Malloch) Malloch 1933 : 241 Malloch 1933 : 241 Stuardo Ortiz 1946 : 149 Frey 1960 : 28 Soós 1968 : 2 Apiochaeta excepta Lonsdale & Marshall 2008c : 17 Lonsdale & Marshall 2008c : 17 Lonsdale & Marshall 2008c : 17 lenticula Lonsdale & Marshall. Lonsdale & Marshall 2008c : 18 Lonsdale & Marshall 2008c : 18 mallochi Lonsdale & Marshall. Lonsdale & Marshall 2008c : 20 Lonsdale & Marshall 2008c : 20 philippii (Rondani) Peratochetvs Philippii Rondani 1863 : 43 Lonsdale & Marshall 2008c : 20 Schiner 1868 : 236 Wulp 1882 : 90 Handlirsch 1884 : 142 Czerny 1903 : 94 philippii Rondani Czerny 1903 : 94 picta Schiner Alloclusia philippii Melander & Argo 1924 : 32 Malloch 1933 : 235 Stuardo Ortiz 1946 : 149 Frey 1960 : 28 Soós 1968 : 2 Gimnomera Brèthes 1924 : 107 Malloch 1933 : 235 philippii Rondani Apiochaeta philippii Lonsdale & Marshall 2008c : 20 Wulp 1882 : 91 Peratochaetus pictipennis Czerny 1903 : 96 Alloclusia pictipennis Melander & Argo 1924 : 33 Malloch 1933 : 236 Stuardo Ortiz 1946 : 150 Frey 1960 : 28 Soós 1968 : 2 Apiochaeta pictipennis Lonsdale & Marshall 2008c : 21 Malloch 1933 : 237 Malloch 1933 : 237 Stuardo Ortiz 1946 : 150 Frey 1960 : 28 Soós 1968 : 2 Apiochaeta similis Lonsdale & Marshall 2008c : 22 A. similis Malloch 1933 : 238 Malloch 1933 : 238 Frey 1960 : 28 Stuardo Ortiz 1946 : 150 Soós 1968 : 2 Apiochaeta tincticornis Lonsdale & Marshall 2008c : 23 Malloch 1933 : 241 Malloch 1933 : 241 Frey 1960 : 28 Stuardo Ortiz 1946 : 150 Soós 1968 : 2 Malloch 1933 : 242 Stuardo Ortiz 1946 : 150 Lonsdale & Marshall 2008c : 24 varia Malloch Apiochaeta varia Lonsdale & Marshall 2008c : 24 varia var. claripennis Malloch Blanchard 1854 : 452 limbipennis Rondani Rondani 1863 : 43 Lonsdale & Marshall 2008c : 28 vitticollis Blanchard Apiochaeta limbipennis Czerny 1903 : 99 Melander & Argo 1924 : 7 Alloclusia limbipennis Malloch 1933 : 238 Stuardo Ortiz 1946 : 149 Frey 1960 : 28 Soós 1968 : 2 Apiochaeta vitticollis Malloch 1933 : 245 Stuardo Ortiz 1946 : 150 Frey 1960 : 28 Soós 1968 : 2 Lonsdale & Marshall 2008c : 28 World Catalogue of the Druid Flies (Diptera: Schizophora: Clusiidae) Owen Lonsdale Zootaxa 2017 2017-10-12 4333 1 1 85  Lonsdale & Marshall 2008c : 8  Lonsdale & Marshall 2008c : 8  Lonsdale & Marshall 2008c : 9  Lonsdale & Marshall 2008c : 9  Lonsdale & Marshall 2008c : 11  Lonsdale & Marshall 2008c : 12  Lonsdale & Marshall 2008c : 13  Lonsdale & Marshall 2008c : 15  Lonsdale & Marshall 2008c : 15  Lonsdale & Marshall 2008c : 17  Lonsdale & Marshall 2008c : 17  Lonsdale & Marshall 2008c : 18  Lonsdale & Marshall 2008c : 18  Lonsdale & Marshall 2008c : 20  Lonsdale & Marshall 2008c : 20  Lonsdale & Marshall 2008c : 20  Lonsdale & Marshall 2008c : 21  Lonsdale & Marshall 2008c : 22  Lonsdale & Marshall 2008c : 23  Lonsdale & Marshall 2008c : 24  Lonsdale & Marshall 2008c : 28 YMB 574045 Czerny Czerny [238,535,1286,1312] Insecta Clusiidae Apiochaeta Animalia Diptera 46 47 Arthropoda genus     Heterochroa  Schiner 1868: 236. Typespecies,  Heterochroa picta Schiner 1868[=  philippii Rondani 1863], by original designation. Gender feminine. Preoccupied by  HeterochroaBoisduval (Nymphalidae).  Marschall 1873: 333[nomenclator];  Scudder 1884: 147[nomenclator];  Schulze et al. 1930: 1534[nomenclator];  Czerny 1903: 98[as synonym of  Peratochaetus].    Apiochaeta  Czerny 1903: 98[family and genus revision]. Typespecies,  Heterochroa bicolor Schiner 1868, subsequent designation by  Soós 1968: 2. Gender feminine.  Kertész 1906: 321[key];  Melander & Argo 1924: 6[family and genus revision];  Schulze et al. 1926b: 233[nomenclator];  Malloch 1933: 238[genus revision];  Neave 1939a: 251[nomenclator];  Stuardo Ortiz 1946: 150[catalogue, Chile];  Hennig 1952: 22[discussion of immature stages in Clusiidae];  Frey 1960: 28[catalogue, world];  McAlpine 1960: 67[key];  Soós 1964a: 5[key], 1968: 2 [catalogue, Americas south of USA];  Cole & Schlinger 1969: 426[citation];  Lonsdale & Marshall 2008c: 8[genus revision, synonymy of  AlloclusiaHendel, phylogeny], 2010: 1047 [Manual of CentralAmerican Diptera]; Lonsdale et al.2010: 231 [family revision, transferred to Sobarocephalinae from Clusiinae].    Peratochaetus. Misspelling of  PeratochetusRondani[junior synonym of  ClusiaHaliday], and misapplication of name for three species including  PhilippiiRondani,  pictipennisWulpand  ornataJohnson(now Heleomyzidae), with typespecies of  PeratochetusRondani( flava Meigen) treated as  Clusia.  Czerny 1903: 92[synonymy of  HeterochroaSchinerwith  Peratochaetus];  Aldrich 1905: 571[catalogue, North America—only lists ornatusJohnson (currently combined with  SpilochroaWilliston, Heleomyzidae)];  Kertész 1906: 321[key];  Malloch 1933: 234[ Chile, as synonym of  Alloclusia- followed by Frey (1960)and Soós (1964a, 1968)].    Alloclusia  Hendel 1917: 36[as replacement name for  HeterochroaSchiner]. Typespecies,  Peratochetvs philippii Rondani 1863, by original designation. Gender feminine.  Melander & Argo 1924: 32[family and genus revision];  Schulze et al. 1926a: 108[nomenclator];  Malloch 1933: 238[genus revision];  Neave 1939a: 116[nomenclator];  Stuardo Ortiz 1946: 149[catalogue, Chile];  Hennig 1952: 227[discussion of immature stages in Clusiidae];  Frey 1960: 28[catalogue, world];  McAlpine 1960: 67[key];  Soós 1964a: 5[key], 1968: 1 [catalogue, Americas south of USA];  Cole & Schlinger 1969: 426[citation];  Lonsdale & Marshall 2008c: 8[as synonym of  Apiochaeta].   FIGURE 5.Illustrations of Clusiidaefrom Czerny (1903), including  Hendelia beckeriCzerny(figs 1-3),  Apiochaeta philippii(Rondani)(figs 4, 5) and  Apiochaeta bicolor(Schiner)(figs 6, 7).   aterrima(Malloch). NT: Argentina, Chile.    Apiochaeta aterrima  Malloch 1933: 244. Chile. Llanquihue Province: Casa Panque. HT ♂ BMNH.  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA];  Lonsdale & Marshall 2008c: 9[redescription, synonymy of  aequalisMalloch, phylogeny, illustrations of head and male genitalia, map].  Apiochaeta aequalis  Malloch 1933: 245. Argentina. “Bariloche, Rio Negro, Agr.” HT ♀ USNM.  Stuardo Ortiz 1946: 150[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA]; Lonsdale et al.2010: 241 [ lapsus, genes sequenced for phylogenetic analysis];  Lonsdale & Marshall 2008c: 9[as synonym of  aterrimaMalloch].   bicolor(Schiner). NT:Chile    Heterochroa bicolor  Schiner 1868: 237. Chile. Concepción Province: Concepción. HT ♀ NMW.  Handlirsch 1884: 142[discussion of  ClusiaHalidayand  HeterochroaSchiner].    Apiochaeta bicolor.  Czerny 1903: 98[generic combination, illustrations of head and wing];  Melander & Argo 1924: 6[summary of original description];  Malloch 1933: 243[redescription];  Stuardo Ortiz 1946: 150[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA];  Lonsdale & Marshall 2008c: 11[redescription].   chiloensis(Malloch). NT:Chile    Alloclusia chiloensis  Malloch 1933: 238. Chile. Chiloé Province: Isla Chiloé, Castro. HT ♂ BMNH.  Malloch 1933: 238[illustration of male terminalia, photo of wing];  Stuardo Ortiz 1946: 149[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 1[catalogue, Americas south of USA].    Apiochaeta chiloensis.  Lonsdale & Marshall 2008c: 12[generic combination, redescription, phylogeny, illustrations of wing and male genitalia, photo, map, Chile]; Lonsdale et al.2010: 241 [genes sequenced for phylogenetic analysis].    connexaMalloch. NT: Argentina, Chile.     Apiochaeta connexa  Malloch 1933: 244. Argentina. Río Negro Province: Bariloche, Rio Negro. HT ♀ USNM.  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA];  Lonsdale & Marshall 2008c: 13[redescription, phylogeny, map, illustrations of male and female genitalia, Chile], 2010: 1046 [reproduction of illustration of female genitalia].   costalis(Malloch). NT: Argentina, Chile    Alloclusia costalis  Malloch 1933: 239. Argentina. Río Negro Province: Bariloche. HT ♂ USNM.  Malloch 1933: 239[illustration of male terminalia];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA].  Apiochaeta costalis.  Lonsdale & Marshall 2008c: 15[generic combination, redescription, phylogeny, illustration of wing, map, Chile].   dacryodesLonsdale & Marshall. NT:Chile    Apiochaeta dacryodes  Lonsdale & Marshall 2008c: 15. Chile.  Concepción Province: 62 kmS San Pedro. HT ♂ CNC.  Lonsdale & Marshall 2008c: 15[phylogeny, illustrations of male genitalia, map].   excepta(Malloch). NT:Chile.    Alloclusia excepta  Malloch 1933: 241. Chile. Chiloé Province: Isla Chiloé, Ancud. HT ♂ USNM.  Malloch 1933: 241[illustration of male terminalia, photo of wing];  Stuardo Ortiz 1946: 149[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA].    Apiochaeta excepta.  Lonsdale & Marshall 2008c: 17[redescription, generic combination, phylogeny, illustrations of male genitalia, map, Chile]; Lonsdale et al.2010: 241 [genes sequenced for phylogenetic analysis].    furcillataLonsdale & Marshall. NT:Chile.     Apiochaeta furcillata  Lonsdale & Marshall 2008c: 17. Chile.  Concepción Province: 62 kmS San Pedro. HT ♂ CNC.  Lonsdale & Marshall 2008c: 17[phylogeny, illustrations of male genitalia, map].   lenticulaLonsdale & Marshall. NT:Chile.     Apiochaeta lenticula  Lonsdale & Marshall 2008c: 18. Chile. AysénProvince:  RioMañiguales, 72°30′W, 45°25′S. HT ♂ CNC.  Lonsdale & Marshall 2008c: 18[phylogeny, illustrations of wing and male genitalia, map].   mallochiLonsdale & Marshall. NT:Chile.    Apiochaeta mallochi  Lonsdale & Marshall 2008c: 20. Chile.  Concepción Province: 62 kmS San Pedro. HT ♂ CNC.  Lonsdale & Marshall 2008c: 20[phylogeny, map].   philippii(Rondani). NT:Chile.     Peratochetvs Philippii Rondani 1863: 43. Chile. ConcepciónProvince: S Concepción, Laraquete. NT ♂CNC [designated by  Lonsdale & Marshall 2008c: 20].    Heterochroa picta  Schiner 1868: 236. Chile. ST 2♂NMW.  Wulp 1882: 90[redescription];  Handlirsch 1884: 142[redescription, illustration of wing];  Czerny 1903: 94[as synonym of  philippiiRondani].  Peratochaetus Philippii.  Czerny 1903: 94[redescription, synonymy of  pictaSchiner, illustrations of head and wing].  Alloclusia philippii.  Melander & Argo 1924: 32[redescription from literature, generic combination];  Malloch 1933: 235[redescription, illustration of male terminalia, photo of wing];  Stuardo Ortiz 1946: 149[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA].    Gimnomera(?) pinocheti  Brèthes 1924: 107. Chile. Bio Bio. Type information unknown.  Malloch 1933: 235[as synonym of  philippiiRondani].    Apiochaeta philippii.  Lonsdale & Marshall 2008c: 20[generic combination, justified emendation, redescription, phylogeny, illustrations of wing and male genitalia, map, Chile]; Lonsdale et al.2010: 241 [genes sequenced for phylogenetic analysis].    pictipennis(Wulp). NT:Chile.      Heterochroa pictipennis  Wulp 1882: 91. Chile. HT ♀ RMNH.    Peratochaetus pictipennis.  Czerny 1903: 96[quotes description, generic combination].    Alloclusia pictipennis.  Melander & Argo 1924: 33[generic combination, redescription from literature];  Malloch 1933: 236[redescription, illustration of male terminalia, photo of wing];  Stuardo Ortiz 1946: 150[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA].    Apiochaeta pictipennis.  Lonsdale & Marshall 2008c: 21[redescription, generic combination, phylogeny, illustrations of head and male genitalia, map, unnecessary designation of neotype, Chile]; Lonsdale et al.2010: 234 [depository of holotype].    similis(Malloch). NT:Chile.     Alloclusia similis  Malloch 1933: 237. Chile. Llanquihue Province: S Chile, Puerto Varas. HT ♂ BMNH.  Malloch 1933: 237[illustration of male terminalia];  Stuardo Ortiz 1946: 150[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA].    Apiochaeta similis.  Lonsdale & Marshall 2008c: 22[generic combination, redescription, phylogeny, illustrations of male genitalia, photo (♀  A. similis-complex), map, Chile].    tincticornis(Malloch). NT: Argentina, Chile.     Alloclusia tincticornis  Malloch 1933: 238. Chile. Puntra. HT ♂ BMNH.  Malloch 1933: 238[illustration of male terminalia];  Frey 1960: 28[catalogue, world];  Stuardo Ortiz 1946: 150[catalogue, Chile];  Soós 1968: 2[catalogue, Americas south of USA].    Apiochaeta tincticornis.  Lonsdale & Marshall 2008c: 23[generic combination, redescription, phylogeny, illustrations of male genitalia, map, Argentina, Chile].    varia(Malloch). NT:Chile.     Alloclusia varia  Malloch 1933: 241. Chile. Llanquihue Province: Puerto Varas. HT ♂ BMNH.  Malloch 1933: 241[illustration of male terminalia, photo of wing];  Frey 1960: 28[catalogue, world];  Stuardo Ortiz 1946: 150[catalogue, Chile];  Soós 1968: 2[catalogue, Americas south of USA].    Alloclusia varia v. claripennis  Malloch 1933: 242. Chile. Llanquihue Province: Puerto Varas. HT ♂ BMNH.  Stuardo Ortiz 1946: 150[catalogue, Chile];  Lonsdale & Marshall 2008c: 24[as synonym of  variaMalloch].    Apiochaeta varia.  Lonsdale & Marshall 2008c: 24[generic combination, synonymy of  variavar. claripennisMalloch, redescription, phylogeny, illustrations of wing and male genitalia (including annulus), map].    vitticollis(Blanchard). NT:Chile.     Helomyza vitticollis  Blanchard 1854: 452. Chile. NT ♂CNC [same specimen designated as NT ♂  limbipennisRondani], designated by Lonsdale & Marshall (2008c).    Peratochetvs limbipennis  Rondani 1863: 43. Chile. Malcho, Parrel-Talca,  750 m.NT ♂CNC [designated by Lonsdale & Marshall 2008c].  Lonsdale & Marshall 2008c: 28[as synonym of  vitticollisBlanchard].    Apiochaeta limbipennis.  Czerny 1903: 99[generic combination, quotes original description];  Melander & Argo 1924: 7[summary of original description].    Alloclusia limbipennis.  Malloch 1933: 238[redescription, generic combination, illustration of male terminalia, photo of wing];  Stuardo Ortiz 1946: 149[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA]; Lonsdale et al.2010: 241 [ lapsus, genes sequenced for phylogenetic analysis].    Apiochaeta vitticollis.  Malloch 1933: 245[tentative generic combination, quotes original description];  Stuardo Ortiz 1946: 150[catalogue, Chile];  Frey 1960: 28[catalogue, world];  Soós 1968: 2[catalogue, Americas south of USA];  Lonsdale & Marshall 2008c: 28[redescription, phylogeny, illustrations of head, wing and male and female genitalia, photo, map, designation of neotype, Argentina, Chile]. 1671716129 CNC Saint Pierre and Miquelon Concepcion Province 48 49 1 1 1671716124 CNC Saint Pierre and Miquelon Concepcion Province 48 49 1 1 1671716125 [199,992,1501,1523] Chile Aysen 48 49 1 Aysen 1671716120 CNC Chile -45.416668 Rio 1130 -72.5 48 49 1 1 Rio 1671716140 CNC Saint Pierre and Miquelon Concepcion Province 48 49 1 1 1671716170 [199,829,244,266] RMNH Chile Chile 49 50 1 1