Revision of the genus Pelecinobaccha Shannon, description of Relictanum gen. nov., and redescription of Atylobaccha flukiella (Curran, 1941) (Diptera: Syrphidae) Miranda, Gil Felipe Gonçalves Marshall, Stephen A. Skevington, Jeffrey H. Zootaxa 2014 3819 1 1 154 8D88V [151,505,151,177] Insecta Syrphidae Pelecinobaccha GBIF Animalia Diptera 51 52 Arthropoda species manuelorum    Typelocality. Costa Rica, Puntarenas, 4.5 kmNE Progresso, 8°28'N 82°51'W. Holotypefemale INBio.    Male. Head: Black. Face sometimes with lateral traces to whole lateral 1/5 pale, black pilose laterally with golden or white pile on ventral ⅓. Lunule pale, black central macula connected to frontal triangle color by narrow dark vitta. Frontal triangle black pilose, with silvery-white pollen laterally, continuous from the face, and brown pollen medially. Frontal prominence and frontal triangle protuberant. Vertical triangle with black pile in 3 irregular rows; ocellar triangle separated by its length from posterior eye margin. Eye contiguity as long as vertical triangle length. Eye with sub-triangular indentation on posterior margin at level of antenna insertion. Antennal insertions narrowly confluent, ventral margin with long dorsal extension. Dorsal ¼ of occiput with 2 rows of long simple black pile; middle ½ with 2–3 regular anterior rows of simple black pile and 1 regular posterior row of long scalelike white pile; ventral ¼ with 2 irregular rows of scale-like white pile.  Thorax: Scutum black, mostly brown dull pollinose, except white on notopleuron, and with sub-median short pair of vittae of concentrated pale pollen, and pollen sparse on transverse suture; scutum with long erect black pile, except golden to white anterior to scutellum, and pile longer on notopleuron anterior to transverse suture; scutum with continuous anterior row of shining white pile. Scutellum dark brown, with metallic-shining pollen, long black pilose, pile longer than on scutum, sub-scutellar fringe long and golden to white with around 20 pile. Pleuron black, white pilose except black pilose on dorso-posterior of posterior anepisternum (densely arranged), anterior anepimeron, and a few black pile may occur on anterior anepisternum. Plumula long, white. Calypter entirely white. Halter brown, capitulum orange.  Wing: Mostly hyaline with anterior margin diffuse dark brown (cells bc, c, sc, r1, anterior ⅔ of r2+3 and most of r), cell bm slightly dark, entirely microtrichose; alula usually hyaline, large, basally 2 times to apically 4 times as broad as c cell, entirely microtrichose.  Legs: Proleg black, profemur with apical half gradually becoming pale, distinct on inner side, protibia with basal half pale, black pilose. Mesoleg black, apical 1/6 of mesofemur and basal ⅔ of mesotibia pale, mesocoxa with only 1 row of longer pile antero-apically, bare to sparse pile ventrally on mesotrochanter, mesofemur bare ventrally and with irregular rows of long pile laterally. Metaleg black, pale on apex of metafemur, slightly less than the basal ½ of the metatibia and from the apical ½ of the metabasitarsomere to the other tarsomeres, apical tarsomere slightly darker, black pilose, white on metacoxa and on white regions.  Abdomen: Black; ~3 times as long as thorax. First tergite short with rounded lateral lobes, with long pile, black dorso-laterally and white ventro-laterally, with a row of white pile dorso-medially and sparse to bare medially. Second tergite long, slightly constricted medially, length ~3.5 times minimum width, and with rectangular dull black pollinose region medially, and mostly black pilose, except white basally, pile long and erect laterally, appressed dorsally; 2nd sternite with long, erect white pile on basal ½. Third tergite trapezoidal, long and widening apically, length ~2.5 times minimum width, with baso-lateral vittate pale maculae, maculae extending ½ the length of the tergite; 3rd tergite dull black-pollinose except for baso-lateral corners and apex, and with appressed black pile, except slightly longer, erect and white on baso-lateral corners. Fourth tergite sub-quadrate, with basolateral pale triangles, pollinose as 3rd tergite, and with appressed black pile, except white on pale corners. Fifth tergite short, rectangular and wide, with appressed black pile. Sternites black, well sclerotized except for some segments with slightly transparent basal regions. Genitalia: Epandrium notched posteriorly, densely microtrichose. Cercus with 1 to 2 regular rows of pile on medial margin, 3 irregular rows of pile on lateral margin, densely microtrichose. Surstylus directed ventrally, with 5 irregular rows of setulae on ventral, slightly concave side, long pilose on basal ½ of dorsal surface. Subepandrial sclerite weakly sclerotized, with a pair of posteriorly directed short processes. Hypandrium with ventral notch rounded and extending to anterior ⅔. Distiphallus smooth, phallapodeme tapering dorso-ventrally on basal ⅓. Postgonite with sparse pilosity except on apex, pile longer basoventrally, expanded ventro-basally and slightly concave towards the apex on the ventral surface; postgonite apex rounded ventrally, convex anteriorly and acute dorsally.  Female: Like male except: Face distinctly pale on lateral ⅓, more extensively pale pilose; ocellar triangle ~2 times its length from posterior eye margin and ~1 occellus-width from lateral eye margin. Anterior row of pile shorter than posterior one on dorsal ¼ of occiput. Wing with dark anterior margin well delimited and not diffuse. Profemur sometimes entirely pale, protibia sometimes with basal ⅔ pale; 2nd to 4th protarsomeres and 3rd and 4th mesotarsomeres slightly swollen. Metafemur sometimes light brown on basal ½, metatibia with only basal ⅓ pale; metacoxa mostly with black pile. Second abdominal tergite shorter, lateral margin narrowly pale basally and wider on the apical ½. Third abdominal tergite shorter, with baso-lateral pale triangles. Fourth abdominal tergite wider than long, with baso-lateral fasciate pale maculae. Sixth segment conical, slightly longer than 5th tergite. Genitalia: Strongly sclerotized. Seventh tergite rectangular normal, apex convex and slightly extended, basal extensions slightly shorter than 6th segment, with pair of short latero-ventral extensions, extensive membranous region between 7th and 8th segments. Eighth tergite unsclerotized medially appearing as a pair of separate sclerites. Tenth tergite as a pair of sclerites fused to the dorsal surface of the cerci. Cercus with 1 row of pile on apical margin.  Length. 9.5–12.5mm; wing 7.5–10.5mm.   Distribution. Costa Rica(Puntarenas).  Primary type. Holotype: Female in good condition. Left proleg and genitalia dissected and placed in individual plastic tubes pinned with the specimen. Holotypelabels: ‘ COSTA RICA, Prov. Puntarenas, // Send. Fila Palmital Chiquisa, 4.5 // km NE. de Progresso. 1600m. 16 // JUL 1997. E. Navarro. // L_S_319950_598950 #47425’ ‘ Holotype//  Pelecinobaccha//  manuelorum// Miranda’ [red label].   Paratypes. COSTA RICA. Puntarenas, Est. Las Alturas, 1 kmNorte de Las Alturas, 1500m, L_S_322700_591400, #6024, 28–30 Sep 1994, M. A. Zumbado ( 1 ♂, INBIO CRI002 494374); ..., Monteverde Biol. Res., 1500m, cloud forest, 13 Jun 2000, M. Buck (1 ♀, DEBU00118179); ..., San Luis, Fca. Buen Amigo Monteverde, 4 kmS de la reserva, 1350m, con Red., L_N_ 250850_449250, #474754, Jul 1997, Z. Fuentes ( 1 ♂, INBIO CRI002 560229); ..., Send. Fila Palmital Chichiza, 4.5 kmNE de Progresso, 1600m, L_S_319950_598950, #47692, 13 Aug 1997, A. Picado ( 2 ♂, INBIO CRI002 559279 & 559285); ..., San Vito de Coto, Las Alturas Biol. Sta., 8°57'N 82°58'W, 15–1700m, 12–14 Aug 1995, S. A. Marshall(1 ♀, DEBU); ..., 1500m, 14 Aug 1995, J.R. Vockeroth ( 1 ♂, CNC Diptera 160777).   Etymology.The specific epithet is a patronym honouring Manuel Zumbado and Manuel Solis, colleagues and friends. It is to be treated as a noun in the genitive case.