Ogloblin 1936 : 38 Gonatocerus ( Gonatocerus ) grandis (Ogloblin) Gonatocerus grandis (Ogloblin) Yoshimoto 1990 : 40 Luft Albarracin et al. 2009 : 9 Lymaenon grandis Ogloblin Loiácono et al. 2005 : 17 Review of Gonatocerus (Hymenoptera: Mymaridae) in the Neotropical region, with description of eleven new species Triapitsyn, Serguei V. Huber, John T. Logarzo, Guillermo A. Berezovskiy, Vladimir V. Aquino, Daniel A. Zootaxa 2010 2010-05-14 2456 1 243 (Ogloblin, 1936) Ogloblin 1936 [151,838,1483,1509] Insecta Mymaridae Gonatocerus Animalia Hymenoptera 156 157 Arthropoda species grandis Cosmocomoidea     Lymaenon grandis  Ogloblin 1936: 38–41 + plates (láminas) II and III (illustrations). Type locality: Loreto, Misiones, Argentina.   Gonatocerus( Gonatocerus) grandis(Ogloblin): De Santis 1967: 104 (catalog).    Gonatocerus grandis(Ogloblin):  Yoshimoto 1990: 40(list);  Luft Albarracin et al.2009: 9(list; distribution in Argentina).    Lymaenon grandisOgloblin:  Loiácono et al.2005: 17(information on potential typespecimens).     Typematerial examined. Lectotypefemale [ MLPA], here designated to avoid confusion about the status of the typespecimens of this species, on slide labeled: “  Gonatocerus grandisA. O ♀ Loreto,  Misiones  2.iii.1932A. A. O.”. The lectotype, although rather poorly cleared, is in relatively good condition except it lacks almost the entire flagellum of the left antenna except for F1, is well spread out and mounted dorsoventrally.  Paralectotypes[all MLPA]: 1 ♂on slide labeled: “  Lymaenon grandis[A. O] ♂ Loreto,  Misiones.  XI.1933 A. O.”;  1 ♂on slide labeled: “  Lymaenon grandis[ A. O.] ♂ Loreto,  Misiones  15.VI.1934.”; 1 ♂ on slide labeled: “  Gonatocerus grandisA. O.  ♂ Loreto,  Misiones  27.viii.1934 Selva.”. Thesefour specimens definitely belonged to the syntypeseries of  Lymaenon grandisbecause they match the collecting dates given in the original description, even though it is evident that the females were listed under the males and vice versa. Ogloblinalso apparently miscalculated the number of the specimens in the syntypeseries, which appears to consist of 1 femaleand 5 males(of which 2 maleshave matching published collecting dates), and not of 3 femalesand 2 malesas mistakenly given in the original description. Thelabel data of these two additional males [both MLPA] are as follows: 1 ♂ on slide labeled: 1. “  Gonatocerus grandisA. O.  ♂ Loreto. Mis.  2.vi.1932A. A. O.  Typus! Fig. Ant.[ip, iR]”; 2. “3856”. Thismale is designated as a paralectotypeas there is very little doubt that it was part of the syntypeseries because the drawing of the male antenna given by Ogloblin (1936, Plate III, fig. 24)was made from this specimen according to the inscription in pencil in Russianon the label. Also 1 ♂(a very probable paralectotype) on slide labeled: “  Gonatocerus grandisA. O ♂  25.iv.1932. A. O. Loreto, Misiones.”.   FIGURES 321–323.  Gonatocerus( Cosmocomoidea) gracilicornis♂ (Loreto, Misiones, Argentina): 321, antenna; 322, wings; 323, genitalia.   FIGURES 324, 325.  Gonatocerus( Cosmocomoidea) grandis♀ (lectotype): 324, antenna; 325, wings.    Materialexamined. ARGENTINA. MISIONES, San Ignacio, Chacra Yabebirí( Yabebiry),  17.iii.1952, A.A. Ogloblin[ 1 ♀, MLPA].   BRAZIL. SANTA CATARINA, Nova Teutonia, 27°11’S 52°23’W,  11.vii.1937, F. Plaumann[ 1 ♀, BMNH].   Redescription.FEMALE ( lectotypeand non-type specimens). Body length 1890–2478µm. Body and appendages mostly light brown except ocelli, eyes, trabeculae, 4 bands on gastral terga, and metatibia brown; flagellum dark brown. Antenna ( Figs 324, 326) with radicle 4.3–4.6x as long as wide, about 0.3x total length of scape, rest of scape 3.7–3.8x as long as wide, scape longitudinally striate; pedicel much shorter than F1; F1 the longest funicle segment, at least a little longer than F2, and with 1 mps; F2 slightly longer than following funicle segments but F2–F4 more or less subequal in length and a little longer than F5–F7 (F6 about as long as F7 and a little shorter than F5), F8 the shortest funicle segment and about as long as wide or just slightly wider than long, F2–F8 each with 2 mps; clava large, with 8 mps, 3.9–4.2x as long as wide, about as long as combined length of F4–F8. Mesosoma. Pronotum very short medially, 2-lobed (mediolongitudinally divided), not entire as drawn (for the male) by Ogloblin (1936). Propodeum ( Fig. 327) with well-developed submedian carinae and several transverse rugosities in posterior half between submedian and lateral carinae; submedian carinae meeting anteriorly and extending to anterior margin of propodeum. Forewing ( Figs 325, 328) 3.2–3.3x as long as wide; longest marginal seta 0.14–0.17x maximum wing width; disc at least slightly infumate, a little more so behind stigmal vein, bare behind venation except for a few setae behind stigmal vein and apex of marginal vein. Hind wing ( Figs 325, 328) 15–17xas long as wide; disc mostly bare except for rows of setae along margins and a few scattered setae basally and apically, slightly infumate; longest marginal seta 1.2–1.4x maximum wing width. Metasoma about as long as mesosoma. Petiole 1.3–1.4x as long as wide, notably narrower basally than distally. Ovipositor about 0.75–0.8x length of gaster, slightly exserted beyond gastral apex by about 0.08x own length; ovipositor length: mesotibia length ratio 1.1–1.2:1. Measurements (µm) of the lectotype. Total body length: 2478; head 344; mesosoma 1015; petiole 123; gaster 1033; ovipositor 806. Antenna: radicle 197; rest of scape 430; pedicel 88; F1 136; F2 115; F3 112; F4 109; F5 103; F6 94; F7 91; F8 64; clava 497. Forewing 2210:689; longest marginal seta 106. Hind wing 1660:112; longest marginal seta 137.   FIGURES 326–328.  Gonatocerus( Cosmocomoidea) grandis♀ (Nova Teutonia, Santa Catarina, Brazil): 326, antenna; 327, dorsellum and propodeum; 328, wings. MALE ( paralectotypesand the probable paralectotype). Body length 1770–2370 µm. Similar to female except for normal sexually dimorphic features and the following. Body generally darker, often with most of median areas of mesoscutum, scutellum and dorsellum ( Fig. 329) brown, as described by Ogloblin (1936). Antenna with scape 2.7–2.8x as long as wide; all flagellomeres longer than scape. Forewing 2.8–3.2x as long as wide. Genitalia as in Fig. 330.   Diagnosis.  Gonatocerus( Cosmocomoidea) grandisis a member of the  morrillisubgroup of the aterspecies group. It is a large species (body length about 1.9–2.5 mm) and females are characterized by their peculiar antenna ( Figs 324, 326) with F1 the longest funicle segment and bearing 1 mps, F2–F8 each with 2 mps, and the large clava about as long as the combined length of F4–F8.   Distribution.NEOTROPICAL: Argentina, and Brazil*.   Hosts.Unknown. 1639354320 1932-03-02 MLPA Misiones & A.A. Ogloblin & The Loreto 156 157 1 1 Loreto lectotype 1639354361 1933-11 MLPA Misiones. & A.A. Ogloblin Loreto 156 157 2 2 Loreto paralectotype 1639354329 1934-06-15 A.A. Ogloblin & Misiones 157 158 Loreto 156 157 2 2 Loreto paralectotype 1639354328 1934-08-27 MLPA Selva. & These & Ogloblin & The Misiones 157 158 14 4 10 Loreto syntype 1639354334 [1038,1431,431,458] 1932-06-02 Loreto 157 158 1 1 Loreto syntype 1639354343 1932-04-25 Russian & Also & A. O. Loreto This Fig. Ant. 157 158 2 2 Loreto paralectotype 1639354356 1952-03-17 MLPA A. A. Ogloblin Argentina Material Chacra Yabebiri San Ignacio 158 159 1 1 MISIONES 1639354346 1937-07-11 BMNH F. Plaumann Brazil -27.183332 Nova Teutonia 1238 -52.383335 158 159 1 1 SANTA CATARINA