Brachymeria Chalcis minuta Thaumatelia Chalcis separata Brachymeria Oncochalcis Oncochalcis marginata Brachymeria Holochalcis Holochalcis madagascariensis Brachymeria Tumidicoxa Tumidicoxa nigra Brachymeria Thaumatelia Chalcis separata Brachymeria Thaumateliana Thaumateliana bicolor Thaumatelia Pseudepitelia Pseudepitelia rubrifemur Brachymeria Brachepitelia Brachepitelia rubripes Brachymeria Tumidicoxoides Tumidicoxoides kurandaensis Brachymeria Tumidicoxella Tumidicoxa Tumidicoxa (Tumidicoxella) nigra Microchalcis Microchalcis atricorpus Dirrhinomorpha Dirrhinomorpha angusta Brachymeria Brachymeria Meyeriella Meyeriella indica Brachymeria Neobrachymeria Brachymeria Brachymeria confalonierii Australochalcis Australochalcis humilicrus Brachymeria Matsumurameria Brachymeria Chalcis taiwanus Gahanula Brachymeria discreta Brachymeria Revision of the family Chalcididae (Hymenoptera, Chalcidoidea) from Vietnam, with the description of 13 new species Narendran, T. C. van Achterberg, Cornelis ZooKeys 2016 576 1 202 62GP4 92426 Westwood, 1829 Westwood 1829 Insecta Chalcididae Brachymeria CoL Animalia Brachymeria Hymenoptera 18 19 Arthropoda genus  Taxon classification Animalia Hymenoptera Chalcididae    BrachymeriaWestwood (in Stephens), 1829: 36. Type species: Chalcis minutaFabricius; designated by Westwood, 1839.  ThaumateliaKirby, 1883: 60. Type species: Chalcis separataWalker, by monotypy (synonymised with BrachymeriaWestwood by Halstead 1991).  OncochalcisCameron, 1904: 162. Type species: Oncochalcis marginataCameron, by monotypy; (synonymised with BrachymeriaWestwood by  Nikol'skaya1960).  HolochalcisKieffer, 1905: 258. Type species: Holochalcis madagascariensisKieffer, by subsequent designation of Gahan and Fagan (1923); (synonymised with BrachymeriaWestwood by Narendran (in Subba Rao & Hayat), 1987b).  TumidicoxaGirault, 1911: 378. Type species: Tumidicoxa nigraGirault; by original designation; (synonymised with BrachymeriaWestwood by Girault (1913)).  ThaumateliaKirby, 1883: 60. Type species Chalcis separataWalker, by monotypy; (synonymised with BrachymeriaWestwood by Halstead 1991).  ThaumatelianaGirault, 1912: 160-161. Type species: Thaumateliana bicolorGirault, by monotypy; (synonymised with ThaumateliaKirby by Narendran and Verghese 1989).  PseudepiteliaGirault, 1913: 104. Type species: Pseudepitelia rubrifemurGirault, by original designation; (synonymised with BrachymeriaWestwood by Girault 1915a).  BrachepiteliaGirault, 1913a: 106. Type species: Brachepitelia rubripesGirault, by original designation and monotypy; (synonymised with BrachymeriaWestwood by Girault (1915a)).  TumidicoxoidesGirault, 1913b: 86. Type species: Tumidicoxoides kurandaensisGirault, by original designation; (synonymy with BrachymeriaWestwood by Girault (1926)).  TumidicoxellaGirault, 1913c: 74. As a subgenus of Tumidicoxa; type species: Tumidicoxa (Tumidicoxella) nigraGirault by original designation.  MicrochalcisGirault, 1915a: 328. Type species: Microchalcis atricorpusGirault by original designation; (synonymised by Boucek, 1988).  DirrhinomorphaGirault & Dodd (in Girault 1915a): 327. Type species: Dirrhinomorpha angustaGirault & Dodd, by original designation; (synonymised with BrachymeriaWestwood and treated as subgenus of Brachymeriaby Boucek1988).  MeyeriellaKrausse, 1917: 95. Type species: Meyeriella indicaKrausse, by monotypy (synonymised with BrachymeriaWestwood by Narendran, 1986).  NeobrachymeriaMasi, 1929c: 196-198. As a subgenus of Brachymeria. Type species: Brachymeria confalonieriiMasi by original designation.  AustralochalcisGirault, 1939: 326. Type species: Australochalcis humilicrusGirault, by original designation and monotypy (listed as synonym of BrachymeriaWestwood by Boucek, 1988).  MatsumurameriaHabu, 1960: 209. As a subgenus of Brachymeria. Type species: Chalcis taiwanusMatsumura, by original designation.  GahanulaBurks, 1960: 261. Type species: Brachymeria discretaGahan, by original designation (as a subgenus of Brachymeria).  Diagnosis. Head oval in profile; scrobe deep with carinate margins; in some species head with pre-orbital or post-orbital carinae or with both carinae present; malar sulcus carinate or ridged; antennal formula 11171 (clava 1 to 3 segmented). Mesosoma with umbilicate punctures; fore wing with PMV usually half or about half as long as MV and usually twice as long as STV. Hind coxa in ♀ in some cases with an inner ventro-mesal tooth; hind femur with a ventral row of irregular teeth and in some species with an inner basal tooth; hind tibia arcuate; metasoma sessile, T1 always the longest; ovipositor sheath slightly compressed and slightly exserted; in some species metasoma elongate.  Hosts. The species are mostly primary parasitoids in pupae of holometabolous insects, especially of Lepidoptera, but some species attack Diptera, Coleopteraor Hymenoptera. Most species are primary parasitoids, but some are hyperparasitoids attacking parasitoid Hymenopteraor Dipterain Lepidoptera.  Distribution. Worldwide.  Key to Vietnamese species of BrachymeriaWestwood     Brachymeria taiwanaHabu  2   3  4   Brachymeria podagrica(Fabricius)   Brachymeria minuta(Linnaeus)  5  6  746  Brachymeria lasus(Walker)  3333  Brachymeria coxodentataJoseph, Narendran & Joy  7  10   Brachymeria carinataJoseph, Narendran & Joy  8   Brachymeria margaroniaeJoseph, Narendran & Joy  9  7068  Brachymeria shansiensisHabu  3838  Brachymeria excarinataGahan   Brachymeria scutellocarinataJoseph, Narendran & Joy  or 11  12  14  2827  Brachymeria aureaGirault   2629 13   Brachymeria megaspila(Cameron)  5253  Brachymeria marmonti(Girault)   Brachymeria bengalensis(Cameron)  15   Brachymeria semirusulasp. n.  16  17  18  36  Brachymeria euploeaeWestwood)  42  Brachymeria jambolanaGahan   Brachymeria lugubris(Walker)  19  4445  Brachymeria kamijoiHabu  404856 20  25  Brachymeria alternipes(Walker)  404856 21   4150 22  23  49  Brachymeria longiscaposaJoseph, Narendran & Joy   Brachymeria himeHabu   Brachymeria olethria(Waterston)  24  57  Brachymeria neowiebesinasp. n.   Brachymeria wiebesinaJoseph, Narendran & Joy  Figure 25. Brachymeria alternipes(Walker), ♂, Vietnam, Hoang Lien N. R., habitus, lateral.  Figure 26. Brachymeria alternipes(Walker), ♂, Vietnam, Hoang Lien N. R., mesonotum, dorsal.  Figures 27-28. Brachymeria aurea(Girault), ♀, Vietnam, Cat TienN. P. 27 habitus lateral 28 mesonotum dorsal.  Figures 29-30. Brachymeria bengalensis(Cameron), ♀, Vietnam, Nui ChuaN. P. 29 habitus lateral 30 scutellum dorsal.  Figures 31-32. Brachymeria carinataJoseph, Narendran & Joy, ♀, Vietnam, Nui ChuaN. P. 31 habitus lateral 32 scutellum, dorsal.  Figures 33-35. Brachymeria coxodentataJoseph, Narendran & Joy, ♀, Vietnam, Cat Ba N. P. 33 habitus lateral 34 scutellum dorsal 35 hind coxa ventral.  Figures 36-37. Brachymeria euploeae(Westwood), ♀, Vietnam, Chu Yang Sin N. P. 36 habitus lateral 37 scutellum dorsal.  Figures 38-39. Brachymeria excarinataGahan, ♀, Vietnam, Cat TienN. P. 38 habitus lateral 39 scutellum dorsal.  Figures 40-41. Brachymeria himeHabu, ♀, Vietnam, Cat TienN. P. 40 habitus lateral 41 scutellum dorsal.  Figures 42-43. Brachymeria jambolanaGahan, ♀, Vietnam, Cat TienN. P. 42 habitus lateral 43 mesonotum dorsal.  Figures 44-45. Brachymeria kamijoiHabu, ♀, Vietnam, Chu Yang Sin N. P. 44 habitus lateral 45 mesonotum dorsal.  Figures 46-47. Brachymeria lasus(Walker), ♀, Vietnam, Cat TienN. P. 46 habitus lateral 47 mesonotum dorsal.  Figures 48-50. Brachymeria longiscaposaJoseph, Narendran & Joy, ♀, Vietnam, Cat TienN. P. 48 habitus lateral 49 head anterior 50 mesonotum dorsal.  Figure 51. Brachymeria lugubris(Walker), ♀, Vietnam, Cat TienN. P., habitus lateral.  Figures 52-53. Brachymeria marmonti(Girault), ♀, Vietnam, Cat TienN. P. 52 habitus lateral 53 scutellum dorsal.  Figures 54-55. Brachymeria minuta(Linnaeus), ♀, Vietnam, Nui ChuaN. P. 54 habitus lateral 55 mesonotum dorsal.  Figure 56. Brachymeria neowiebesinasp. n., ♀, holotype, habitus lateral.  Figures 57-58. Brachymeria neowiebesinasp. n., ♀, holotype. 57 mesonotum dorsal 58 head anterior.  Figure 59. Brachymeria podagrica(Fabricius), ♀, Vietnam, Nui ChuaN. P., habitus lateral.  Figures 60-61. Brachymeria podagrica(Fabricius), ♀, Vietnam, Nui ChuaN. P. 60 head anterior 61 mesosoma dorsal.  Figure 62. Brachymeria scutellocarinataJoseph, Narendran & Joy, ♀, Vietnam, Cat TienN. P., habitus lateral.  Figures 63-64. Brachymeria scutellocarinataJoseph, Narendran & Joy, ♀, Vietnam, Cat TienN. P. 63 mesonotum dorsal 64 head anterior.  Figure 65. Brachymeria semirusulasp. n., ♀, holotype, habitus lateral.  Figures 66-67. Brachymeria semirusulasp. n., ♀, holotype. 66 mesonotum dorsal 67 head anterior.  Figures 68-70. Brachymeria shansiensisHabu, ♀, Nui ChuaN. P. 68 habitus lateral 69 scutellum dorsal 70 hind tarsus dorsal.