Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains
Wang, XIN-PING
Griswold, CHARLES E.
Miller, JEREMY A.
Zootaxa
2010
2593
1
127
37KG6
Wang
Wang
Arachnida
Agelenidae
Draconarius
CoL
Animalia
Araneae
31
35
Arthropoda
species
dubius
(Figs 99-120, 537)
Draconarius dubius Wang2003: 530, figs 29A-D ( Holotype male, maleand female paratypesfrom Lushui County, Yunnan, China, in NHUand CAS, examined). Wang & Jaeger2008: 2279, fig. 4. FIGURES93-98. Draconarius disgregus Wang2003, females (93-94, 97-98) ( HNUCASENT9023091) and female (95-96) ( HNUCASENT9022203) from Lushui County, Pianma Township, Chan Yan He. 93-96. Epigynum (ventral, dorsal); 97. Eyes, fronto-dorsal; 98. Labium, ventral ( L/ W=1.08). Additional material examined. CHINA: Yunnan: Lushui County: 1♀, Feng Xue Yakou, on Pianma Road, N25.97288°, E98.68336°, 3150 m, hand collecting under rocks, May 11, 2005, C. Griswold& D. Kavanaugh( HNU, CASENT9022195); 3♂, Feng Xue Yakou, 100 mS of Pianma Road, N25.97288°, E98.68336°, 3150 m, Rhododendron /Bamboo thicket, 25 pitfall traps, May 11-21, 2005, C. Griswold, D. Kavanaugh& K.J. Guo( HNU, CASENT9022107); 2♂, Feng Xue Yakou, 100 mS of Pianma Road, N25.97288°, E98.68336°, 3150 m, Rhododendron/Bamboo thicket, pitfall traps, May 11-21, 2005, C. Griswold, D. Kavanaugh& K.J. Guo( CAS, CASENT9030744); 1♀, Feng Xue Yakou, E side on Pianma Road, N25.97288°, E98.68336°, 3150 m, hand collecting under rocks, May 17, 2005, C. Griswold( HNU, CASENT9022252); 2♀, Pianma Township, Feng Xue Yakou, N25.97814°, E98.67508°, 3088 m, roadside, May 19, 2005, K.J. Guo( CAS, CASENT9022041); 1♀, Pianma Township, Feng Xue Yakou, N26.00949°, E98.61704°, 3142 m, along the road, May 17, 2005, K.J. Guo( HNU, CASENT9022072). Fugong County: 1♀, Lishadi, Shibali, N27.16650°, E98.77936°, 2537 m, forest, webs on tree trunks, at night, August 5, 2005, P. Paquin, D. Kavanaugh( HNU, CASENT9022570); 1♀, Lishadi, Shibali Yaku (pass #31), N27.21230°, E98.69600°, 3604 m, boulder field to bamboo thicket, under rocks, August 5, 2005, P. Paquin( HNU, CASENT9022561); 1♀, Gaoligongshan, 7.61 kmdirectly WNN of Shibali, Rhododendron thicket, turning rocks amid bamboo, N27.20662°, E98.71773°, 3292 m, May 6, 2004, C. Griswold( CAS, CASENT 9020783); 1♀, Gaoligongshan, 1 kmE Yaping Pass, turning rocks among dormant bamboo, N27.20854°, E98.20854°, 3506 m, May 6, 2004, C. Griswold( CAS, CASENT9020765); 1♀, Yaping subdistrict, Gaoligongshan, Maxidi, 7.2 kmabove Shibali, N27.16784°, E98.77782°, 3059 m, May 4, 2004, H.M. Yan, G.X. Peng( CAS, CASENT9020581); 1♀, Lishadi subdistrict, Gaoligongshan, 12 kmabove Shibali on Yaping Road, N27.20658°, E98.21782°, 3100-3280 m, May 8, 2004, H.M. Yan( CAS, CASENT9020600); 1♀, Gaoligongshan, 7.41 kmWNW of Shibali, 36 kmNNW of Fugong, beneath objects amidst dormant bamboo, along snowfield and avalanche debris, N27.20629°, E98.72001°, 3336 m, May 8, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9019948); 1♀, Gaoligongshan, Shibali Forest Station, in moist stream gully, N27.16894°, E98.77205°, 2650 m, May 9, 2004, C. Griswold& H.M. Yan( CAS, CASENT9019910); 2♀, Gaoligongshan, Shibali Forest Station, in moist stream gully, N27.16894°, E98.77205°, 2650m, May 9, 2004, C. Griswold& H.M. Yan( HNU, CASENT9019909); 1♀, Lishadi, Shibali Yaku (pass #31), N27.21230°, E98.69600°, 3604 m, boulder field to bamboo thicket, under rocks, August 5, 2005, P. Paquin( HNU, CASENT9022562); 1♀, Lishadi, 1 kmbefore Shibali Yaku (pass #31), 900 mN of the road, N27.21447°, E98.70064°, 3585 m, August 12, 2005, P. Paquin( HNU, CASENT9022556); 2♀, Lishadi, 500 mW of Shibali, N27.16650°, E98.77936°, 2537 m, deciduous forest litter, August 4, 2005, P. Paquin( HNU, CASENT9022576); 1♀, Lishadi, Shibali, N27.16650°, E98.77936°, 2537 m, forest, webs on tree trunks, at night, August 5, 2005, P. Paquin, D. Kavanaugh( HNU, CASENT9022571); 1♀, Lishadi, 500 mbefore Shibali Yaku (pass #31), N27.21354°, E98.70021°, 3585 m, stable scree slope on soil, August 7, 2005, P. Paquin( HNU, CASENT9022550); 1♀, Lishadi, 500 mW of Shibali, N27.16650°, E98.77936°, 2537 m, deciduous forest litter, August 4, 2005, P. Paquin( HNU, CASENT9022577); 1♀, Gaoligongshan, 1 kmE Yaping Pass turning rocks among dormant bamboo, N27.20854°, E98.20854°, 3506 m, understory of good forest on E-facing slope; night collecting, May 6, 2005, C. Griswold( CAS, CASENT9020767); 1♀, Lishadi, 10.5 kmW of Shibali, 100 mN road, N27.20192°, E98.71321°, 3250 m, dead conifer logs in conifer forest with Rhododendron patch, August 17, 2005, P. Paquin( CAS, CASENT9022565); 1♀, Lishadi, Shibali Yaku (pass #31), N27.21230°, E98.69600°, 3604 m, boulder field to bamboo thicket, under rocks, August 5, 2005, P. Paquin( CAS, CASENT9022560); 1♀, Lishadi, 10 kmW of Shibali, N27.20055°, E98.71399°, 3221 m, mature pine forest with bamboo understory, under rocks & logs, August 6, 2005, P. Paquin( CAS, CASENT9022538); 2♀, Gaoligongshan, 1 kmE Yaping Pass, turning rocksamong dormant bamboo, N27.20854°, E98.20854°, 3506 m, May 6, 2004, C. Griswold( CAS, CASENT9020766); 1♀, LiShadi subdist, Gaoligongshan, 12 kmabove Shibali on Yaping Road, N27.20658°, E98.21782°, 3100-3280 m, May 8, 2004, H.M. Yan( CAS, CASENT9020601); 1♀, Gaoligongshan, 10.1-11.5 road km from Shibali on Yaping Road, N27.20049°, E98.71354°to N27.20676°, E98.71763, 3225-3290 m, night collectingalong road, May 8, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020035); 1♀, Lishadi subdistrict, Gaoligongshan, Maxidi, 11.5 kmabove Shibali on Yaping Road, N27.20490°, E98.71480°, 3266 m, May 6, 2004, H.M. Yan( CAS, CASENT9025795); 1♀, Gaoligongshan, 7.61 kmdirectly WNN of Shibali, Rhododendron thicket, turning rocks amid bamboo, N27.20662°, E98.71773°, 3292 m, May 6, 1998, C. Griswold( HNU, CASENT9020784); 3♀, Lishadi Township, 8.5 kmabove Shibali on Shibali Road, North Fork of Yamu He, N27.18416°, E98.72026°, 3100 m, May 9, 2004, D. Kavanaugh& H.B. Liang( HNU, CASENT9023030); 1♀, Gaoligongshan, 1 kmE Yaping Pass turning rocksamong dormant bamboo, N27.20854°, E98. 20854°, 3506 m, May 6, 2004, C. Griswold( HNU, ENT 9020768); 1♀, Pihe Township, Yueliangtian Village, N26.56784°, E098.90884°, 1520 m, August 20/23/ 24, 2005, G. Tang( CAS, CASENT9025591); 4♀, Lishadi Township, Yakou of Shibali, N27.21234°, E098.69601°, 3615 m, August 5/7, 2005, G. Tang( CAS, CASENT9025583). Gongshan County: 1♀, Cikai Township, 41 kmW of Gongshan on Dulong Valley Road at Dabadi, N27.79655°, E098.50562°, 3000 m, September 27 to October 6, 2002, D. Kavanaugh, P.E. Marek, D.Z. Dong, & X.C. Liang( CAS, CASENT9025832,); 1♀, Cikai Township, 53 kmW of Gongshan on Dulong Valley Road, N27.77422°, E098.44716°, 3380m, October 1, 2002, D. Kavanaughcollector ( CAS, CASENT9025855); 1♀, Cikai Township, Dabadi to Yakou, tunnel, marshland, N27°27', E98°37', 3000 m, October 5, 2002, H.M. Yan( CAS, CASENT9025858); 1♀, Dulongjiang Township, near San Dui, N27°43'3.6'', E98°25'17'', 2620 m, October 29, 2004, V.F. Lee( HNU, CASENT9023065); 1♀, Dulongjiang Township, near San Dui, N27°43'3.6'', E98°25'17'', 2620 m, October 29, 2004, V.F. Lee( CAS, CASENT9023066).
Diagnosis: The female is similar to D. patellabifidus Wang 2003 inlacking epigynal teeth and having broad spermathecae, but can be recognized by the absence of atrial ridges and the posteriorly broad, anteriorly less broad, diverging spermathecae (Figs 104, 109-120). The male can be easily recognized by the long patellar apophysis (longer than patellar length), the simple median apophysis, the bifurcate conductor, and the prolaterally originating embolus (Figs 99-101).
Description: See Wang (2003). Photos of habitus, eyes, labium and genitalia are provided in this study (Figs 99-120).
Distribution: China (Yunnan: Lushui, Fugong, Gongshan) (Fig. 537).
923902927
HNU
China
Lushui County
1
1
Yunnan
holotype
923902926
HNU
China
Lushui County
Yunnan
paratype
923902938
CAS
China
Lushui County
Yunnan
holotype
923902939
2005-05-11
hand collecting
HNU
C. Griswold, D. Kavanaugh
China
Lushui County
3150 m
25.97288
Feng Xue Yakou, on Pianma Road
0
98.68336
CASENT 9022195
1
1
Yunnan
Other material
923902946
2005-05-11
2005-05-21
2005-05-11
25 pitfall traps
HNU
C. Griswold, D. Kavanaugh, K. J. Guo
China
Lushui County
3150 m
25.97288
Feng Xue Yakou, 100 m S of Pianma Road
0
98.68336
CASENT 9022107
3
3
Yunnan
Other material
923902940
2005-05-11
2005-05-21
2005-05-11
pitfall traps
CAS
C. Griswold, D. Kavanaugh, K. J. Guo
China
Lushui County
3150 m
25.97288
Feng Xue Yakou, 100 m S of Pianma Road
0
98.68336
CASENT 9030744
2
2
Yunnan
Other material
923902933
2005-05-17
hand collecting
HNU
C. Griswold
China
Lushui County
3150 m
25.97288
Feng Xue Yakou, E side on Pianma Road
0
98.68336
CASENT 9022252
1
1
Yunnan
Other material
923902925
2005-05-19
CAS
K. J. Guo
China
Lushui County
3088 m
25.97814
Pianma Township, Feng Xue Yakou
0
98.67508
CASENT 9022041
2
2
Yunnan
Other material
923902936
2005-05-17
HNU
K. J. Guo
China
Lushui County
3142 m
26.00949
Pianma Township, Feng Xue Yakou
0
98.61704
CASENT 9022072
1
1
Yunnan
Other material
923902930
2005-08-05
at night
HNU
P. Paquin, D. Kavanaugh
China
Fugong County
2537 m
27.1665
Lishadi, Shibali
0
98.77936
CASENT 9022570
1
1
Yunnan
Other material
923902929
2005-08-05
HNU
P. Paquin
China
Fugong County
3604 m
27.2123
Lishadi, Shibali Yaku (pass 31)
0
98.696
CASENT 9022561
1
1
Yunnan
Other material
923902941
2004-05-06
turning rocks amid bamboo
CAS
C. Griswold
China
Fugong County
3292 m
27.20662
Gaoligongshan, 7.61 km directly WNN of Shibali
0
98.71773
CASENT 9020783
1
1
Yunnan
Other material
923902928
2004-05-06
turning rocks among dormant bamboo
CAS
C. Griswold
China
Fugong County
3506 m
27.20854
Gaoligongshan, 1 km E Yaping Pass
0
98.20854
CASENT 9020765
1
1
Yunnan
Other material
923902932
2004-05-04
CAS
H. M. Yan, G. X. Peng
China
Fugong County
3059 m
27.16784
Yaping subdistrict, Gaoligongshan, Maxidi, 7.2 km above Shibali
0
98.77782
CASENT 9020581
1
1
Yunnan
Other material
923902934
2004-05-08
CAS
H. M. Yan
China
Fugong County
3100 - 3280 m
27.20658
Lishadi subdistrict, Gaoligongshan, 12 km above Shibali on Yaping Road
0
98.21782
CASENT 9020600
1
1
Yunnan
Other material
923902945
2004-05-08
CAS
C. Griswold, D. Kavanaugh
China
Fugong County
3336 m
27.20629
Gaoligongshan, 7.41 km WNW of Shibali, 36 km NNW of Fugong
0
98.72001
CASENT 9019948
1
1
Yunnan
Other material
923902964
2004-05-09
CAS
C. Griswold, H. M. Yan
China
Fugong County
2650 m
27.16894
Gaoligongshan, Shibali Forest Station
0
98.77205
CASENT 9019910
1
1
Yunnan
Other material
923902935
2004-05-09
HNU
C. Griswold, H. M. Yan
China
Fugong County
2650 m
27.16894
Gaoligongshan, Shibali Forest Station
0
98.77205
CASENT 9019909
2
2
Yunnan
Other material
923902931
2005-08-05
under rocks
HNU
P. Paquin
China
Fugong County
3604 m
27.2123
Lishadi, Shibali Yaku (pass 31)
0
98.696
CASENT 9022562
1
1
Yunnan
Other material
923902943
2005-08-12
HNU
P. Paquin
China
Fugong County
3585 m
27.21447
Lishadi, 1 km before Shibali Yaku (pass 31), 900 m N of the road
0
98.70064
CASENT 9022556
1
1
Yunnan
Other material
923902942
2005-08-04
HNU
P. Paquin
China
Fugong County
2537 m
27.1665
Lishadi, 500 m W of Shibali
0
98.77936
CASENT 9022576
2
2
Yunnan
Other material
923902944
2005-08-05
at night
HNU
P. Paquin, D. Kavanaugh
China
Fugong County
2537 m
27.1665
Lishadi, Shibali
0
98.77936
CASENT 9022571
1
1
Yunnan
Other material
923902948
2005-08-07
HNU
P. Paquin
China
Fugong County
3585 m
27.21354
Lishadi, 500 m before Shibali Yaku (pass 31)
0
98.70021
CASENT 9022550
1
1
Yunnan
Other material
923902951
2005-08-04
HNU
P. Paquin
China
Fugong County
2537 m
27.1665
Lishadi, 500 m W of Shibali
0
98.77936
CASENT 9022577
1
1
Yunnan
Other material
923902947
2005-05-06
turning rocks among dormant bamboo, night collecting
CAS
C. Griswold
China
Fugong County
3506 m
27.20854
Gaoligongshan, 1 km E Yaping Pass
0
98.20854
CASENT 9020767
1
1
Yunnan
Other material
923902952
2005-08-17
CAS
P. Paquin
China
Fugong County
3250 m
27.20192
Lishadi, 10.5 km W of Shibali, 100 m N road
0
98.71321
CASENT 9022565
1
1
Yunnan
Other material
923902949
2005-08-05
CAS
P. Paquin
China
Fugong County
3604 m
27.2123
Lishadi, Shibali Yaku (pass 31)
0
98.696
CASENT 9022560
1
1
Yunnan
Other material
923902959
2005-08-06
CAS
P. Paquin
China
Fugong County
3221 m
27.20055
Lishadi, 10 km W of Shibali
0
98.71399
CASENT 9022538
1
1
Yunnan
Other material
923902962
2004-05-06
turning rocks
CAS
C. Griswold
China
Fugong County
3506 m
27.20854
Gaoligongshan, 1 km E Yaping Pass
0
98.20854
CASENT 9020766
2
2
Yunnan
Other material
923902965
2004-05-08
CAS
H. M. Yan
China
Fugong County
3100 m
27.20658
LiShadi subdist, Gaoligongshan, 12 km above Shibali on Yaping Road
0
98.21782
CASENT 9020601
1
1
Yunnan
Other material
923902966
2004-05-08
night collecting
CAS
C. Griswold, D. Kavanaugh
China
Fugong County
3258 m
27.20676
Gaoligongshan, 10.1 - 11.5 road km from Shibali on Yaping Road
0
98.71354
CASENT 9020035
1
1
Yunnan
Other material
923902961
2004-05-06
CAS
H. M. Yan
China
Fugong County
3266 m
27.2049
Lishadi subdistrict, Gaoligongshan, Maxidi, 11.5 km above Shibali on Yaping Road
0
98.7148
CASENT 9025795
1
1
Yunnan
Other material
923902950
1998-05-06
turning rocks amid bamboo
HNU
C. Griswold
China
Fugong County
3292 m
27.20662
Gaoligongshan, 7.61 km directly WNN of Shibali
0
98.71773
CASENT 9020784
1
1
Yunnan
Other material
923902960
2004-05-09
HNU
D. Kavanaugh, H. B. Liang
China
Fugong County
3100 m
27.18416
Lishadi Township, 8.5 km above Shibali on Shibali Road, North Fork of Yamu He
0
98.72026
CASENT 9023030
3
3
Yunnan
Other material
923902954
2004-05-06
turning rocks
HNU
C. Griswold
China
Fugong County
3506 m
27.20854
Gaoligongshan, 1 km E Yaping Pass
98.20854
ENT 9020768
1
1
Yunnan
Other material
923902953
2005-08-20
2005-08-24
2005-08-20
CAS
G. Tang
China
Fugong County
1520 m
26.56784
Pihe Township, Yueliangtian Village
0
98.90884
CASENT 9025591
1
1
Yunnan
Other material
923902970
2005-08-05
2005-08-07
2005-08-05
CAS
G. Tang
China
Fugong County
3615 m
27.21234
Lishadi Township, Yakou of Shibali
0
98.69601
CASENT 9025583
4
4
Yunnan
Other material
923902956
2002-09-27
2002-10-06
2002-09-27
CAS
D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang
China
Gongshan County
3000 m
27.79655
Cikai Township, 41 km W of Gongshan on Dulong Valley Road at Dabadi
0
98.50562
CASENT 9025832
1
1
Yunnan
Other material
923902958
2002-10-01
CAS
D. Kavanaugh
China
Gongshan County
3380 m
27.77422
Cikai Township, 53 km W of Gongshan on Dulong Valley Road
0
98.44716
CASENT 9025855
1
1
Yunnan
Other material
923902957
2002-10-05
CAS
H. M. Yan
China
Gongshan County
3000 m
27.45
Cikai Township, Dabadi to Yakou
1
98.61667
CASENT 9025858
1
1
Yunnan
Other material
923902968
2004-10-29
HNU
V. F. Lee
China
Gongshan County
2620 m
27.717667
Dulongjiang Township, near San Dui
1
98.42139
CASENT 9023065
1
1
Yunnan
Other material
923902989
2004-10-29
CAS
V. F. Lee
China
Gongshan County
2620 m
27.717667
Dulongjiang Township, near San Dui
1
98.42139
CASENT 9023066
1
1
Yunnan
Other material