Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains Wang, XIN-PING Griswold, CHARLES E. Miller, JEREMY A. Zootaxa 2010 2593 1 127 37KG6 Wang Wang Arachnida Agelenidae Draconarius CoL Animalia Araneae 31 35 Arthropoda species dubius  (Figs 99-120, 537)    Draconarius dubius Wang2003: 530, figs 29A-D ( Holotype male,  maleand female paratypesfrom  Lushui County, Yunnan, China, in NHUand  CAS, examined). Wang & Jaeger2008: 2279, fig. 4.   FIGURES93-98. Draconarius disgregus Wang2003, females (93-94, 97-98) ( HNUCASENT9023091) and female (95-96) ( HNUCASENT9022203) from Lushui County, Pianma Township, Chan Yan He. 93-96. Epigynum (ventral, dorsal); 97. Eyes, fronto-dorsal; 98. Labium, ventral ( L/ W=1.08).  Additional material examined. CHINA: Yunnan: Lushui County: 1♀, Feng Xue Yakou, on Pianma Road, N25.97288°, E98.68336°,  3150 m, hand collecting under rocks,  May 11, 2005, C. Griswold& D. Kavanaugh( HNU, CASENT9022195);  3♂, Feng Xue Yakou, 100 mS of Pianma Road, N25.97288°, E98.68336°,  3150 m, Rhododendron /Bamboo thicket, 25 pitfall traps,  May 11-21, 2005, C. Griswold, D. Kavanaugh& K.J. Guo( HNU, CASENT9022107);  2♂, Feng Xue Yakou, 100 mS of Pianma Road, N25.97288°, E98.68336°,  3150 m, Rhododendron/Bamboo thicket, pitfall traps,  May 11-21, 2005, C. Griswold, D. Kavanaugh& K.J. Guo( CAS, CASENT9030744);  1♀, Feng Xue Yakou, E side on Pianma Road, N25.97288°, E98.68336°,  3150 m, hand collecting under rocks,  May 17, 2005, C. Griswold( HNU, CASENT9022252);  2♀, Pianma Township, Feng Xue Yakou, N25.97814°, E98.67508°,  3088 m, roadside,  May 19, 2005, K.J. Guo( CAS, CASENT9022041);  1♀, Pianma Township, Feng Xue Yakou, N26.00949°, E98.61704°,  3142 m, along the road,  May 17, 2005, K.J. Guo( HNU, CASENT9022072).  Fugong County: 1♀, Lishadi, Shibali, N27.16650°, E98.77936°,  2537 m, forest, webs on tree trunks, at night,  August 5, 2005, P. Paquin, D. Kavanaugh( HNU, CASENT9022570);  1♀, Lishadi, Shibali Yaku (pass #31), N27.21230°, E98.69600°,  3604 m, boulder field to bamboo thicket, under rocks,  August 5, 2005, P. Paquin( HNU, CASENT9022561);  1♀, Gaoligongshan, 7.61 kmdirectly WNN of Shibali, Rhododendron thicket, turning rocks amid bamboo, N27.20662°, E98.71773°,  3292 m,  May 6, 2004, C. Griswold( CAS, CASENT 9020783);  1♀, Gaoligongshan, 1 kmE Yaping Pass, turning rocks among dormant bamboo, N27.20854°, E98.20854°,  3506 m,  May 6, 2004, C. Griswold( CAS, CASENT9020765);  1♀, Yaping subdistrict, Gaoligongshan, Maxidi, 7.2 kmabove Shibali, N27.16784°, E98.77782°,  3059 m,  May 4, 2004, H.M. Yan, G.X. Peng( CAS, CASENT9020581);  1♀, Lishadi subdistrict, Gaoligongshan, 12 kmabove Shibali on Yaping Road, N27.20658°, E98.21782°,  3100-3280 m,  May 8, 2004, H.M. Yan( CAS, CASENT9020600);  1♀, Gaoligongshan, 7.41 kmWNW of Shibali, 36 kmNNW of Fugong, beneath objects amidst dormant bamboo, along snowfield and avalanche debris, N27.20629°, E98.72001°,  3336 m,  May 8, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9019948);  1♀, Gaoligongshan, Shibali Forest Station, in moist stream gully, N27.16894°,  E98.77205°,  2650 m,  May 9, 2004, C. Griswold& H.M. Yan( CAS, CASENT9019910);  2♀, Gaoligongshan, Shibali Forest Station, in moist stream gully, N27.16894°, E98.77205°,  2650m,  May 9, 2004, C. Griswold& H.M. Yan( HNU, CASENT9019909);  1♀, Lishadi, Shibali Yaku (pass #31), N27.21230°, E98.69600°,  3604 m, boulder field to bamboo thicket, under rocks,  August 5, 2005, P. Paquin( HNU, CASENT9022562);  1♀, Lishadi, 1 kmbefore Shibali Yaku (pass #31), 900 mN of the road, N27.21447°, E98.70064°,  3585 m,  August 12, 2005, P. Paquin( HNU, CASENT9022556);  2♀, Lishadi, 500 mW of Shibali, N27.16650°, E98.77936°,  2537 m, deciduous forest litter,  August 4, 2005, P. Paquin( HNU, CASENT9022576);  1♀, Lishadi, Shibali, N27.16650°, E98.77936°,  2537 m, forest, webs on tree trunks, at night,  August 5, 2005, P. Paquin, D. Kavanaugh( HNU, CASENT9022571);  1♀, Lishadi, 500 mbefore Shibali Yaku (pass #31), N27.21354°, E98.70021°,  3585 m, stable scree slope on soil,  August 7, 2005, P. Paquin( HNU, CASENT9022550);  1♀, Lishadi, 500 mW of Shibali, N27.16650°, E98.77936°,  2537 m, deciduous forest litter,  August 4, 2005, P. Paquin( HNU, CASENT9022577);  1♀, Gaoligongshan, 1 kmE Yaping Pass turning rocks among dormant bamboo, N27.20854°, E98.20854°,  3506 m, understory of good forest on E-facing slope; night collecting,  May 6, 2005, C. Griswold( CAS, CASENT9020767);  1♀, Lishadi, 10.5 kmW of Shibali, 100 mN road, N27.20192°, E98.71321°,  3250 m, dead conifer logs in conifer forest with Rhododendron patch,  August 17, 2005, P. Paquin( CAS, CASENT9022565);  1♀, Lishadi, Shibali Yaku (pass #31), N27.21230°, E98.69600°,  3604 m, boulder field to bamboo thicket, under rocks,  August 5, 2005, P. Paquin( CAS, CASENT9022560);  1♀, Lishadi, 10 kmW of Shibali, N27.20055°, E98.71399°,  3221 m, mature pine forest with bamboo understory, under rocks & logs,  August 6, 2005, P. Paquin( CAS, CASENT9022538);  2♀, Gaoligongshan, 1 kmE Yaping Pass, turning rocksamong dormant bamboo, N27.20854°, E98.20854°,  3506 m,  May 6, 2004, C. Griswold( CAS, CASENT9020766);  1♀, LiShadi subdist, Gaoligongshan, 12 kmabove Shibali on Yaping Road, N27.20658°, E98.21782°,  3100-3280 m,  May 8, 2004, H.M. Yan( CAS, CASENT9020601);  1♀, Gaoligongshan, 10.1-11.5 road km from Shibali on Yaping Road, N27.20049°, E98.71354°to N27.20676°, E98.71763,  3225-3290 m, night collectingalong road,  May 8, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020035);  1♀, Lishadi subdistrict, Gaoligongshan, Maxidi, 11.5 kmabove Shibali on Yaping Road, N27.20490°, E98.71480°,  3266 m,  May 6, 2004, H.M. Yan( CAS, CASENT9025795);  1♀, Gaoligongshan, 7.61 kmdirectly WNN of Shibali, Rhododendron thicket, turning rocks amid bamboo, N27.20662°, E98.71773°,  3292 m,  May 6, 1998, C. Griswold( HNU, CASENT9020784);  3♀, Lishadi Township, 8.5 kmabove Shibali on Shibali Road, North Fork of Yamu He, N27.18416°, E98.72026°,  3100 m,  May 9, 2004, D. Kavanaugh& H.B. Liang( HNU, CASENT9023030);  1♀, Gaoligongshan, 1 kmE Yaping Pass turning rocksamong dormant bamboo, N27.20854°, E98. 20854°,  3506 m,  May 6, 2004, C. Griswold( HNU, ENT 9020768);  1♀, Pihe Township, Yueliangtian Village, N26.56784°, E098.90884°,  1520 m,  August 20/23/ 24, 2005, G. Tang( CAS, CASENT9025591);  4♀, Lishadi Township, Yakou of Shibali, N27.21234°, E098.69601°,  3615 m,  August 5/7, 2005, G. Tang( CAS, CASENT9025583).  Gongshan County: 1♀, Cikai Township, 41 kmW of Gongshan on Dulong Valley Road at Dabadi, N27.79655°, E098.50562°,  3000 m,  September 27 to October 6, 2002, D. Kavanaugh, P.E. Marek, D.Z. Dong, & X.C. Liang( CAS, CASENT9025832,);  1♀, Cikai Township, 53 kmW of Gongshan on Dulong Valley Road, N27.77422°, E098.44716°,  3380m,  October 1, 2002, D. Kavanaughcollector ( CAS, CASENT9025855);  1♀, Cikai Township, Dabadi to Yakou, tunnel, marshland, N27°27', E98°37',  3000 m,  October 5, 2002, H.M. Yan( CAS, CASENT9025858);  1♀, Dulongjiang Township, near San Dui, N27°43'3.6'', E98°25'17'',  2620 m,  October 29, 2004, V.F. Lee( HNU, CASENT9023065);  1♀, Dulongjiang Township, near San Dui, N27°43'3.6'', E98°25'17'',  2620 m,  October 29, 2004, V.F. Lee( CAS, CASENT9023066).  Diagnosis: The female is similar to D. patellabifidus Wang 2003 inlacking epigynal teeth and having broad spermathecae, but can be recognized by the absence of atrial ridges and the posteriorly broad, anteriorly less broad, diverging spermathecae (Figs 104, 109-120). The male can be easily recognized by the long patellar apophysis (longer than patellar length), the simple median apophysis, the bifurcate conductor, and the prolaterally originating embolus (Figs 99-101).   Description: See Wang (2003). Photos of habitus, eyes, labium and genitalia are provided in this study (Figs 99-120).  Distribution: China (Yunnan: Lushui, Fugong, Gongshan) (Fig. 537). 923902927 HNU China Lushui County 1 1 Yunnan holotype 923902926 HNU China Lushui County Yunnan paratype 923902938 CAS China Lushui County Yunnan holotype 923902939 2005-05-11 hand collecting HNU C. Griswold, D. Kavanaugh China Lushui County 3150 m 25.97288 Feng Xue Yakou, on Pianma Road 0 98.68336 CASENT 9022195 1 1 Yunnan Other material 923902946 2005-05-11 2005-05-21 2005-05-11 25 pitfall traps HNU C. Griswold, D. Kavanaugh, K. J. Guo China Lushui County 3150 m 25.97288 Feng Xue Yakou, 100 m S of Pianma Road 0 98.68336 CASENT 9022107 3 3 Yunnan Other material 923902940 2005-05-11 2005-05-21 2005-05-11 pitfall traps CAS C. Griswold, D. Kavanaugh, K. J. Guo China Lushui County 3150 m 25.97288 Feng Xue Yakou, 100 m S of Pianma Road 0 98.68336 CASENT 9030744 2 2 Yunnan Other material 923902933 2005-05-17 hand collecting HNU C. Griswold China Lushui County 3150 m 25.97288 Feng Xue Yakou, E side on Pianma Road 0 98.68336 CASENT 9022252 1 1 Yunnan Other material 923902925 2005-05-19 CAS K. J. Guo China Lushui County 3088 m 25.97814 Pianma Township, Feng Xue Yakou 0 98.67508 CASENT 9022041 2 2 Yunnan Other material 923902936 2005-05-17 HNU K. J. Guo China Lushui County 3142 m 26.00949 Pianma Township, Feng Xue Yakou 0 98.61704 CASENT 9022072 1 1 Yunnan Other material 923902930 2005-08-05 at night HNU P. Paquin, D. Kavanaugh China Fugong County 2537 m 27.1665 Lishadi, Shibali 0 98.77936 CASENT 9022570 1 1 Yunnan Other material 923902929 2005-08-05 HNU P. Paquin China Fugong County 3604 m 27.2123 Lishadi, Shibali Yaku (pass 31) 0 98.696 CASENT 9022561 1 1 Yunnan Other material 923902941 2004-05-06 turning rocks amid bamboo CAS C. Griswold China Fugong County 3292 m 27.20662 Gaoligongshan, 7.61 km directly WNN of Shibali 0 98.71773 CASENT 9020783 1 1 Yunnan Other material 923902928 2004-05-06 turning rocks among dormant bamboo CAS C. Griswold China Fugong County 3506 m 27.20854 Gaoligongshan, 1 km E Yaping Pass 0 98.20854 CASENT 9020765 1 1 Yunnan Other material 923902932 2004-05-04 CAS H. M. Yan, G. X. Peng China Fugong County 3059 m 27.16784 Yaping subdistrict, Gaoligongshan, Maxidi, 7.2 km above Shibali 0 98.77782 CASENT 9020581 1 1 Yunnan Other material 923902934 2004-05-08 CAS H. M. Yan China Fugong County 3100 - 3280 m 27.20658 Lishadi subdistrict, Gaoligongshan, 12 km above Shibali on Yaping Road 0 98.21782 CASENT 9020600 1 1 Yunnan Other material 923902945 2004-05-08 CAS C. Griswold, D. Kavanaugh China Fugong County 3336 m 27.20629 Gaoligongshan, 7.41 km WNW of Shibali, 36 km NNW of Fugong 0 98.72001 CASENT 9019948 1 1 Yunnan Other material 923902964 2004-05-09 CAS C. Griswold, H. M. Yan China Fugong County 2650 m 27.16894 Gaoligongshan, Shibali Forest Station 0 98.77205 CASENT 9019910 1 1 Yunnan Other material 923902935 2004-05-09 HNU C. Griswold, H. M. Yan China Fugong County 2650 m 27.16894 Gaoligongshan, Shibali Forest Station 0 98.77205 CASENT 9019909 2 2 Yunnan Other material 923902931 2005-08-05 under rocks HNU P. Paquin China Fugong County 3604 m 27.2123 Lishadi, Shibali Yaku (pass 31) 0 98.696 CASENT 9022562 1 1 Yunnan Other material 923902943 2005-08-12 HNU P. Paquin China Fugong County 3585 m 27.21447 Lishadi, 1 km before Shibali Yaku (pass 31), 900 m N of the road 0 98.70064 CASENT 9022556 1 1 Yunnan Other material 923902942 2005-08-04 HNU P. Paquin China Fugong County 2537 m 27.1665 Lishadi, 500 m W of Shibali 0 98.77936 CASENT 9022576 2 2 Yunnan Other material 923902944 2005-08-05 at night HNU P. Paquin, D. Kavanaugh China Fugong County 2537 m 27.1665 Lishadi, Shibali 0 98.77936 CASENT 9022571 1 1 Yunnan Other material 923902948 2005-08-07 HNU P. Paquin China Fugong County 3585 m 27.21354 Lishadi, 500 m before Shibali Yaku (pass 31) 0 98.70021 CASENT 9022550 1 1 Yunnan Other material 923902951 2005-08-04 HNU P. Paquin China Fugong County 2537 m 27.1665 Lishadi, 500 m W of Shibali 0 98.77936 CASENT 9022577 1 1 Yunnan Other material 923902947 2005-05-06 turning rocks among dormant bamboo, night collecting CAS C. Griswold China Fugong County 3506 m 27.20854 Gaoligongshan, 1 km E Yaping Pass 0 98.20854 CASENT 9020767 1 1 Yunnan Other material 923902952 2005-08-17 CAS P. Paquin China Fugong County 3250 m 27.20192 Lishadi, 10.5 km W of Shibali, 100 m N road 0 98.71321 CASENT 9022565 1 1 Yunnan Other material 923902949 2005-08-05 CAS P. Paquin China Fugong County 3604 m 27.2123 Lishadi, Shibali Yaku (pass 31) 0 98.696 CASENT 9022560 1 1 Yunnan Other material 923902959 2005-08-06 CAS P. Paquin China Fugong County 3221 m 27.20055 Lishadi, 10 km W of Shibali 0 98.71399 CASENT 9022538 1 1 Yunnan Other material 923902962 2004-05-06 turning rocks CAS C. Griswold China Fugong County 3506 m 27.20854 Gaoligongshan, 1 km E Yaping Pass 0 98.20854 CASENT 9020766 2 2 Yunnan Other material 923902965 2004-05-08 CAS H. M. Yan China Fugong County 3100 m 27.20658 LiShadi subdist, Gaoligongshan, 12 km above Shibali on Yaping Road 0 98.21782 CASENT 9020601 1 1 Yunnan Other material 923902966 2004-05-08 night collecting CAS C. Griswold, D. Kavanaugh China Fugong County 3258 m 27.20676 Gaoligongshan, 10.1 - 11.5 road km from Shibali on Yaping Road 0 98.71354 CASENT 9020035 1 1 Yunnan Other material 923902961 2004-05-06 CAS H. M. Yan China Fugong County 3266 m 27.2049 Lishadi subdistrict, Gaoligongshan, Maxidi, 11.5 km above Shibali on Yaping Road 0 98.7148 CASENT 9025795 1 1 Yunnan Other material 923902950 1998-05-06 turning rocks amid bamboo HNU C. Griswold China Fugong County 3292 m 27.20662 Gaoligongshan, 7.61 km directly WNN of Shibali 0 98.71773 CASENT 9020784 1 1 Yunnan Other material 923902960 2004-05-09 HNU D. Kavanaugh, H. B. Liang China Fugong County 3100 m 27.18416 Lishadi Township, 8.5 km above Shibali on Shibali Road, North Fork of Yamu He 0 98.72026 CASENT 9023030 3 3 Yunnan Other material 923902954 2004-05-06 turning rocks HNU C. Griswold China Fugong County 3506 m 27.20854 Gaoligongshan, 1 km E Yaping Pass 98.20854 ENT 9020768 1 1 Yunnan Other material 923902953 2005-08-20 2005-08-24 2005-08-20 CAS G. Tang China Fugong County 1520 m 26.56784 Pihe Township, Yueliangtian Village 0 98.90884 CASENT 9025591 1 1 Yunnan Other material 923902970 2005-08-05 2005-08-07 2005-08-05 CAS G. Tang China Fugong County 3615 m 27.21234 Lishadi Township, Yakou of Shibali 0 98.69601 CASENT 9025583 4 4 Yunnan Other material 923902956 2002-09-27 2002-10-06 2002-09-27 CAS D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang China Gongshan County 3000 m 27.79655 Cikai Township, 41 km W of Gongshan on Dulong Valley Road at Dabadi 0 98.50562 CASENT 9025832 1 1 Yunnan Other material 923902958 2002-10-01 CAS D. Kavanaugh China Gongshan County 3380 m 27.77422 Cikai Township, 53 km W of Gongshan on Dulong Valley Road 0 98.44716 CASENT 9025855 1 1 Yunnan Other material 923902957 2002-10-05 CAS H. M. Yan China Gongshan County 3000 m 27.45 Cikai Township, Dabadi to Yakou 1 98.61667 CASENT 9025858 1 1 Yunnan Other material 923902968 2004-10-29 HNU V. F. Lee China Gongshan County 2620 m 27.717667 Dulongjiang Township, near San Dui 1 98.42139 CASENT 9023065 1 1 Yunnan Other material 923902989 2004-10-29 CAS V. F. Lee China Gongshan County 2620 m 27.717667 Dulongjiang Township, near San Dui 1 98.42139 CASENT 9023066 1 1 Yunnan Other material