Ichnea elongata Knull, 1939: 27 Classification, Natural History, and Evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VIII. The Genera Acanthocollum Opitz, Stegnoclava Opitz, and Ichnea Laporte Opitz, Weston The Coleopterists Bulletin 2010 mo 9 2010-12-01 64 1 65 (Klug, 1842) Klug 1842 [799,1101,624,645] Insecta Cleridae Ichnea GBIF Animalia Coleoptera 32 33 Arthropoda species aterrima  ( Figs. 71–74, 124, 125, 139, 167, 189, 193, 194; Map 4)   Enoplium aterrimum Klug, 1842: 378. Holotypemale. México(ZMHB). (Specimen point mounted, gender label affixed to paper point; mount card; locality/identification label; type label; specimen number label; ZMHB repository label; acronymic label; holotypelabel; identification label, plastic vial with aedeagus.) Spinola 1844: 162; Lacordaire 1857: 477; Gemminger and Harold 1869: 1751; Lohde 1900: 100; Schenkling 1903: 102, 1910: 122; Gorham 1883: 181; Knull 1939: 27; Blackwelder 1945: 389; Corporaal 1950: 270; Opitz 2002: 273, 275 (misspelled as atterima).   Ichnea mexicana Thomson, 1860: 65. Holotype ♀and four paratypes. Mexico. (MNHN).  Newsynonymy. Thecharacteristics used to describe this nominal species represent intraspecific variation. Gemmingerand Harold1869: 1751; Chevrolat1876: 36; Lohde1900: 101; Gorham1883: 179; Schenkling1910: 123; Corporaal1950: 271; Opitz1997: 75 ( Fig. 56, not 55 which is a photograph of  I. praeusta).   Ichnea religiosa Chevrolat, 1874: 323. Lectotype ♂. Herein designated. México, Oaxaca, Da Salle(MNHN). New synonymy. The characteristics used to describe this nominal species represent intraspecific variation. Gorham 1883: 180; Lohde 1900: 102; Schenkling 1903: 102, 1910: 123; Blackwelder 1945: 389; Corporaal 1950: 272.   Ichnea vitticollis Kuwert, 1894:11. Lectotype, gender unknown. Hereindesignated. México(MNHN). New synonymy. The characteristics used to describe this nominal species represent intraspecific variation. Lohde 1900: 102; Schenkling 1903: 102, 1910: 124; Blackwelder 1945: 389; Corporaal 1950: 271.   Ichnea kuwertiLohde, 1899: 305. New synonymy. Unnecessary replacement name for  Ichnea vitticollisKuwert, which is not congeneric with  Ichnea vitticollisGorham (1877: 415). The characteristics used to describe this nominal species represent intraspecific variation.    Ichnea elongata Knull, 1939: 27. Holotype ♀. Williams, Arizona,  9-VIII-1939, D. J. and J. N. Knull(FMNH).  Newsynonymy. Corporaal1950: 27; Opitz2002: 278.   Diagnosis.This is the only species in the genus that has a phenon whose specimens are entirely dark brown to black. Members of this dark phenon are known only from southwestern U.S.A.and midwestern Mexico. All other members of this species are readily identified by the color of the pronotum whose sides are black and disc center vertically yellow.   Description. Size: Length 7.2 mm; width 2.5 mm. Form( Fig. 139): Narrowly triangular. Integumental color( Figs. 71–74): Dark brown. Head: HW/ PW 1.0; frons indented; EW/ VW 2.6; antenna as in Fig. 167, funicle compressed. Thorax: Pronotum as in Fig. 124, PL/PW 1.1; elytra strongly flared to posterior two-thirds, EL/PL 5.1; EL/EW 2.5; anterior margin of protibia with 12 spines. Male genitalia: Aedeagus as in Fig. 189. Male mesodermal internal reproductive organs( Fig. 193): Medial accessory gland shorter than undulating lateral gland; testis comprised of 12 follicles. Female mesodermal internal reproductive organs( Fig. 194): Spermatheca not capsular; spermathecal gland attached to subapex of spermatheca; bursa copulatrix capsular.  Variation.Size: length 6.0–12.0 mm; width 1.5– 4.0 mm. There is considerable color variation involving the cranium, pronotum, and elytra. Most often the cranium is yellow with postocular dark streaks that blend posteriorly with dark lateral aspects of the pronotum. The latter has a broad, vertical yellow line at mid-disk. Color of the elytra ranges from completely black to completely yellow brown; specimens may have dark and light regions as depicted in Figs. 71–74.  Natural History.Specimens were captured from April through August, at 300–1,800 m, with a Malaise trap, at black light, under bark, and amidst trees in an oak forest. Donald E. Bright collected one specimenin the Chiricahua Mountains of Arizonafrom  Pinus ponderosaC. Lawson (Pinaceae).   Distribution.Known from the USAto Ecuador( Map 4).  Specimens Examined. 205.  USA: New Mexico, Otero Co., Cloudcroft; Grant Co., Pinos Altos:  Arizona, Chiricahua Co, Chiricahua Mountains,  2- VIII-1952, D. J. & J. N. Knull; Coconino Co.,  16 kmSW Flagstaff,  14-VIII-1968, on  Pinus ponderosa, D. E. Bright.   MEXICO: Tamaulipas,  10 kmE Tula,  10-V-1994, 1189 m, oak forest, J. E. Wappes:  Nayarit,  21 kmNW Ahuacation,  25-VIII-1959, R. B. Selander; 16 kmE Tula  10-V-1994, 1188 m Oak Forest, J.E. Wappes: Jalisco, 18 kmN Volcanes, 20°23′N 104°38′W, 29-VI-1995, 1710 m, F. A. Noguera & A. Rodriquez: Veracruz, 10 kmN Fortin, 21-29-VII-1976, E. Giesbert; Cordova (= Cordoba), Salle; Cordoba, 6-VII- 74, 930 m, J. Powell & J. Linsley; Fortin de las Flores, 1-VIII-64, H.V. Daly; 8 kmW Orizaba, 7-VII-1962, J.M. Campbell; 14–16 kmW S’ ntcompn, 4-6-V-1994; 5 kmN Coscomtepec, 7-8-VI-1991, J.A. Chemsak; Estación Biologico Los Tuxtlas, 1-9-VII-1988, J.A. Chemsak: Morelos, 6 kmW Vautepec, 17-18-X-1984, E. Giesbert: Oaxaca, 21 kmS Suchixtepec, 14-VII- 1973, Mastro & Schaffner; idem, 16-VII-1973, Mastro & Schaffner; Yolotepec, Salle: Chiapas, 9.8 kmN Ocosingo, 20-VI-1990, R. Turnbow; 9.6 kmE San Cristóbal de las Casas, 2-VI-1969, H. F. Howden; idem, 19-V-1969, H. F. Howden; Laguna Montebello, 30-VII-1974, collected at night, C. W. O’ Brien; Lagos de Colores, 14-VI- 1969, D. E. Bright; idem, 30-V- 1969, 152 m, Malaise Trap, D. E. Bright; Simojovel, 18-31-VII-1958, J.A. Chemsak.  GUATEMALA: Baja Verapaz: S. Gerónimo, Champion: Chiqimula, 10 kmN Esquipulas, 11-VI-1991, J. E. Wappes; idem, 11- VI-1991, E. Giesbert: Zacapa, 8 kmS San Lorenzo, 22-X-1990, E. Giesbert; 2 kmS San Lorenzo, 3- 5-VI-1989, 1646 m, E. Giesbert: Huehuetenango, 15 kmN Nentón, 5-VI- 1997, 950 m, E. Giesbert & J. Monzon: Izabal, 23 kmSE Morales, 23-27- V-1995, Giesbert & Monzón: Sacatepéquez, Antigua, 20-VIII-1947, 1524 m, C&P Vaurie: El Progreso, Sierra de las Minas, 19 kmN Estanoia de la Virgen, 18-20-IV-1990, 1829 m, E. Giesbert.  BELIZE: Orange Walk, Rio Bravo Cons. area Texas camp, 11-VII-1996, C. W. & L. B. O’ Brien; idem, 13-VII-1996, C. W. & L. B. O’ Brien; idem, 15-VII- 1996, C. W. & L. B. O’ Brien; idem, 18-VII-1996, C. W. & L. B. O’ Brien; idem, 19-VII-1996, C. W. & L. B. O’ Brien.  HONDURAS: Cortes, 1 kmW El Jaral Finca Fe, 8-VIII-1977, C. W. O’ Brien; Peña Blanca, Lago Yojoa, 21-VI-1979, at light, J. A. Chemsak: Atlántida, Parque Nacional Pico Bonito, El Manchon, 15°29′ N18″ 87°07′39″W, 20-III- 2000, A. Lopez Martinez; Tela Tela, 27-V-1993, R. Cave: Yoro, Linda Vista, Pico Pijol, 15°10′N 87°05′W, 13-VII-1996, R. D. Cave; Palo de Comba, 15°11′N 87°39′W, 5-VIII-1995, R. Cave: Olancho, La Muralla Parque Nacional, 24-27-V- 1995, J. E. Wappes; La Muralla Parque Nacional, 24-V-1995, R. Turnbow; idem, 25-V-1995, R. Turnbow; idem, 26-V-1995, R. Turnbow; La Muralla Parque Nacional 14 kmN La Union, 26-27-V-1995, R. D. Cave; La Muralla, 24-V-1995, R. Morris; La Muralla Parque Nacional 15 kmN La Union, 15°07′N 86°45′W, IV-1995, R. Cave; La Muralla Parque Nacional 14 kmN. La Union, 25-26-V-1995, R. D. Cave; La Muralla Parque Nacional, 14 kmN La Unión, 15°07′N 86°45′W, V-1995; 16 kmSW Juticalpa, 16-VI- 1974, 427 m, C. W. O’ Brien: Francisco Morazán, 7.6 kmN Guaimaca, 14°36′N 86°49′W, 26-VI- 1994, 820 m, under bark, J. Ashe & R. Brooks; San Juan del Rancho, Tegucigalpa, 18-X-1987, R. D. Cave; San Antonio de Oriente, El Zamorano, 27-VI-1988, J. Magana: El Paraíso, Cerro Monserrat, 7 kmSW Yuscarán, 8-VI-1994, 1800 m, H. & A. Howden; idem, 30-VI-1994, H. & A. Howden.  NICARAGUA: Matagalpa, Montaña Selva Negra 7.5 kmN Matagalpa, 13°00′1″N 85°54′32″W, 15-16-VI-2001, 1300 m, B. Ratcliffe, M. L. Jameson & R. Cave; Santa Martha, 8 kmN Matagalpa, 12-IX-1976, Edward S. Ross: Granada: Reserva Nacional, Volcán Mombacho, entrance road, 11°50′ 05″N85°58′83″W, 2-VI- 2002, 750 m, on bark, R. Brooks & Z. Falin, & S. Chatzimanolis; idem, Finca Santa Anna, 21-VI- 1998, 600 m, Malaise trap in organic coffee, J. M. Maes; idem, 16-III-1998, Malaise trap in cloud forest, J. M. Maes; idem, 15-IV-1998, Malaise trap in cloud forest, J. M. Maes; idem, Finca El Progresso, 31-III- 1998, Malaise trap in nonorganic coffee, J. M. Maes; idem, 30-IV-1998, Malaise trap in nonorganic coffee, J. M. Maes; idem, 2-VI-1998, Malaise trap in nonorganic coffee, J. M. Maes; idem, 21-VI-1998, Malaise trap in nonorganic coffee, J. M. Maes idem, Finca San Joaquin, 21-VI-1998, Malaise trap in nonorganic coffee, J. M. Maes; idem, 30-VI-1998, Malaise trap in nonorganic coffee, J. M. Maes.  COSTA RICA: Guanacaste, Cerro El Hacha, Santa Rosa NP, 22-VI-13-VII- 1985, 300 m, D. Janzen; idem, 14-IX-5-X-1985, D. Janzen; idem, 24- VII-14V-1985, D. Janzen; 3 kmSE R. Naranjo, 14- 16-VI-1993, F. D. Parker; Estación Cacao SW side Volcan Cacao, 1988, 1000–1400 mMalaise trap; Sendero Santa Maria P.N. Rincon de la Vieja E Las Pailas, 7-23-I- 1994, 800 m Malaise trap, E Araya; Estación Maritza lado oeste Volcán Orosi, II- 1992, 600 m Malaise trap; idem, 1988: San José, 27-VI- 1925, Coil. Schmidt; Farm La Caja 8 kmW San José, XI-XII-1924, Hamburg: Cartago, 7 kmNE Pavones, 30-V-1985, J. Doyen; Turriabla, 3-XII- 1972, 600 m, V. O. Becker; idem, 26-VII- 1973, 600 m, V. O. Becker: Puntarenas, Monteverde, 20- V-1985, 1300 m, J. T. Doyden; Monteverde, 26-V- 3-VI-1984, E. Riley & D. Rider & D. LeDoux; Monteverde, 12-16-VI-1978, E. Giesbert; 4 kmNI San Luis de Guacimal, 26-V-1987, E. Giesbert; San Luis (Monteverde), 12-13-V-1996, 1189 m, E. Giesbert; Las Alturas (San Vito), 9-V-1996, 1500 m, E. Giesbert & H. Lezama:  PANAMA: Chiriquí, Hartmann’ s Finca, 18-V-1996; R. Turnbow; Vic. Cerro Punta, 13-14-V-1981, E. Giesbert; Fortuna Dam, 16-V-1993, E. Giesbert; Pacific Slope Volcán, Hacienda las Lagunas, 7-I-1981, 1300 m, G. Ekis; Fortuna, 17-V-1978, C. W. O’ Brien & G. B. Marshall: Panamá, Cerro Jefe, 11-12-V- 1996, 671 m, Wappes & Huether & Morris; Cerro Jefe, 10 kmNE, 9°12′N 79°21′W, 21-IV- 1973, 700 m, Shockwell; Cerro Jefe, 8 kmNE, 9°12′N 79°21′W, 21-IV- 1973, 700 m, Shockwell; 3 kmS Cerro Jefe, 24-VII- 1970, 700 m, H. Hespenheide; Altos De Pacora, 4-10-I, E. Giesbert; idem, 23-IV- 1995, E. Giesbert; Canal Zone, Madden Forrest, 9°05′N 79°37′W, 17-IV-1976, H. P. Stockwell; 10 kmNE Cerro Jefe, 9°12′N 79°21′W, 21-IV- 1973, 700 m, Stockwell; 8 kmNE Cerro Jefe, 9°12′N 79°21′W, 21-IV- 1973, 700 m, Stockwell; Cerro Campana, 8°40′ N 79°56′W, 9- 5-1973, 850 m, H. A. Hespenheide; Cerro Azul, 4 kmbeyond Goofy Lake, 9°12′N 79°23′W, 700 m, 24-VII-1970, H. A. Hespenheide.  VENEZUELA: Aragua, Rancho Grande, 9-VII-1973, 1100 m, J. Salcedo & J. Claviio: Falcón, Cerro Galicia, 22-XI-1971, 1500 m, J. & B. Bechyne; Guaraque, Mesa de Quintero, 6-VII-1979, 2500–3000 m, B. Bechyne.  COLOMBIA: Cauca, Villa Elvira, 1924, Staudinger.  ECUADOR: Carchí, Chical, 0°56′N 78°11′W, 3-   Map 4.Geographic distribution of  Ichneaspp.  10-VII-1983, 1250 m, R. Davidson; idem,  1-10-VII- 1983, R. Davidson; Hojal,  9-VIII-1983, R. Davidson. Specimensare deposited in ALAS, AMNH, BMNH, CASC, CDAE, CHAH, CMNC, CMNH, CNCI, DEIG, EBCC, EMEC, EMUS, FMNH, FSCA, IMLA, INBC, MIZA, JEWC, JNRC, JPHC, LACM, MCZC, MIUP, MNHN, MRAC, RDCC, RFMC, RGCG, RHTC, SEAN, SEMC, STRI, TAMU, TUMC, UMMZ, UCAG, UNSM, USNM, WOPC, ZMAN, and ZMHB.  Notes.  Ichnea kuwertiLohdeis a name established by Lohde (1899) to replace  Ichnea vitticollisKuwert, a presumed junior homonym of  Ichnea vitticollisGorham. However,  Ichnea vitticollisGorhamis actually a junior synonym of  Apolopha reicheiSpinola (Opitz 1998). [922,1090,725,746] ZMHB Mexico Mexico 32 33 2 holotype [877,1069,1044,1065] MNHN Mexico Opitz New Corporaal 32 33 2 1 holotype [873,1085,1283,1304] MNHN Da Salle Mexico Oaxaca 32 33 1 1 Oaxaca lectotype [861,1021,1469,1490] MNHN Mexico Herein 32 33 1 lectotype [328,496,274,295] 1939-08-09 FMNH D. J. and J. N. Knull & New & Corporaal & Opitz Williams 33 34 1 1 Arizona holotype United States of America Otero Co. Grant Co. 33 34 1 New Mexico 1952-08-02 1968-08-14 1952-08-02 D. J. & J. N. Knull United States of America Coconino Co. Chiricahua Mountains Chiricahua Co 33 34 1 Arizona 1994-05-10 J. E. Wappes Mexico 10 km E Tula 33 34 1 Tamaulipas 1959-08-25 1994-05-10 1959-08-25 R. B. Selander Mexico Tula 33 34 1 Nayarit 1983-07-01 1983-08-09 1983-07-01 ALAS, AMNH, BMNH, CASC, CDAE, CHAH, CMNC, CMNH, CNCI, DEIG, EBCC, EMEC, EMUS, FMNH, FSCA, IMLA, INBC, MIZA, JEWC, JNRC, JPHC, LACM, MCZC, MIUP, MNHN, MRAC, RDCC, RFMC, RGCG, RHTC, SEAN, SEMC, STRI, TAMU, TUMC, UMMZ, UCAG, UNSM, USNM, WOPC, ZMAN R. Davidson & Hojal & R. Davidson. Specimens 35 36 1