Acomopterella Review of the Palaearctic Acomopterella Zaitzev (Diptera, Sciaroidea, Mycetophilidae) Kallweit, Uwe ZooKeys 2013 269 11 32 8ZY82 http://species-id.net/wiki/Acomopterella Zaitzev, 1989 Zaitzev 1989 Insecta Mycetophilidae Acomopterella CoL Animalia Acomopterella Diptera 1 12 Arthropoda genus   AcomopterellaZaitzev, 1989: 134 (description); Sevcik& Chandler: 2008: 63 (discussion).  Type species.  Acomopterella arnaudiZaitzev, 1989: 134 (by original designation) = Tetragoneura fallaxSherman, 1925: 20.  Diagnosis. (based on Zaitzev'soriginal genus description, with additions). Body length 4.6-5.9 mm. Head. With 3 ocelli. Mid ocellus slightly in front of the lateral ocelli. Lateral ocelli scarcely to distinctly remote from eye margin. Eyes with short setulae. Face rather flat, bare or setose. Clypeus oval, slightly bulging, setose. Face and clypeus loosely connected. Palpi pentamerous. Antennae 16-segmented, 2-3 times length of the thorax. Length of mesial flagellomeres 3-5 times their own width. Thorax. Scutum with short, close lying acrostichal and long erect dorsocentral bristles. Scutellum with 2 very long medial bristles. Mediotergite, laterotergites and pleura bare. Wing. Transparent. Wing membrane with microtrichia only. C, the apical part of ta, R1, R5, M1, M2, M4 and CuA with macrotrichia. C extending to the tip of R5. R4 present; ta oblique, slightly shorter than stem of fork M1 + M2. Base of M4 on level of base of ta or slightly more proximal. Vein A well developed, its tip behind the level of base of ta or ending slightly before. Between A and the stem of fork M4 + CuA lies the weakly sclerotized vein CuP, traceable as fold line. Legs. Long. Fore coxae with numerous long bristles on its frontal side. Apical part of mid coxae with bristles on its lateral and frontal surface. Hind coxae with a row of very long bristles on the hind lateral surface. Mid tibiae with well developed tibial organ on its basal part. Tarsal claws with teeth. Empodia well developed. Abdomen. Segments 1-7 of normal size. Tergite 8 vestigial; Sternite 8 exceeding half length of sternite 7. Tergite 9 rectangular, with deep distal incision. Gonocoxites with apical processes.  Comments. The described genus belongs to the tribe Gnoristiniin the subfamily Sciophilinae. It is similar to Dziedzickia marginata( Dziedzicki 1885), but differing bybare laterotergites and a much longer vein ta. It differs from the Holarctic genus AcomopteraVockeroth, 1980, in the position of Sc (in Acomopterellait joins R1, in AcomopteraC), in the shape of the cell between Rs and R4 (distinctly longer in Acomoptera) and in the strikingly longer ta. The genus is also similar to AustrosynaphaTonnoir, 1929, known from South America, New Zealand and Tasmania. Typical in both genera is the absence of setae on the laterotergites and the presence of a relatively long oblique vein ta. In Austrosynaphavein R4 is absent.