Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae) Génier, François Moretto, Philippe Zootaxa 2017 4248 1 1 110 363WG [151,460,223,248] Insecta Scarabaeidae Digitonthophagus Animalia Coleoptera 73 74 Arthropoda species bonasus  2.  Digitonthophagus uksGénier,  new species Distribution: Afghanistan, India, Iran, Pakistan 3.  Digitonthophagus sahelicusMoretto,  new species Distribution: Burkina Faso, Cameroon, Chad,  Djibouti, Eritrea, Mali, Mauritania,  Niger, Senegal, Sudan