Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)
Génier, François
Moretto, Philippe
Zootaxa
2017
4248
1
1
110
363WL
Genier
Genier
[151,561,1186,1213]
Insecta
Scarabaeidae
Digitonthophagus
Animalia
Coleoptera
32
33
Arthropoda
species
falciger
sp. nov.
Typelocality. Forêt de Boulon( 10°16’27”N 04°27’15”W), 270 m, Comoé, Burkina Faso.
Description. Holotype ♂( Fig. 12). Measurements. Length 11.0 mm, width 7.0 mm. Head( Figs. 33–34). Anterior clypeal edge straight on median fourth in dorsal view; clypeofrontal carina broadly arcuate and interrupted at gena. Vertex lacking median tubercle, surface with punctures fine to small, scabrous anteriorly, separated by one to four diameters. Horns moderately long, parallel on apical half in frontal view, gradually tapering from base to apex; posterointernal edge unmodified basally, apicointernal surface lacking granules; genal edge upturned and distinctly angulate on anterior third, forming a very broad angle with clypeal edge. Pronotum( Fig. 60). Surface with granulate punctures restricted to anterior half medially, with distinct, scattered, umbilicate punctures on posterior half of disc; punctures weaker but distinct posterolaterally, with more-or-less distinct, minute punctures throughout. Anteromedian tubercle atrophied, simply round in lateral view, median longitudinal sulcus well defined; surface behind the eyes with a simple, round depression, surface of anterior angles flat; anterior half of lateral edge arcuate in dorsal and lateral view; posterior angles simply arcuate in dorsal view. Anterior hypomeral ridge arcuate anteriorly, nearly straight posteriorly, anterior hypomeral depression surface slightly darker in color medially. Elytra. ( Fig. 12). Intervals 2 and 4 with few scattered fine granules from base to apex. Legs. Protibial apicointernal tooth enlarged, with dorsal ridge extending to apex. Aedeagus( Fig. 124). Parameres with dorsal and ventral edges slightly diverging toward apex in lateral view. Internal sac sclerites( Figs. 150–152). Axial sclerite weekly sclerotized, as long as subaxial sclerite and almost straight. Subaxial sclerite, more-or-less even in width, extending straight and interrupted before apex of frontolateral peripheral sclerite apical portion, with short to long villi on apical sixth. Frontolateral peripheral sclerite basoventral apophysis moderately developed; with a spiniform medioventral carina; right lateral fold produced into a bifurcate process apically; left lateral lobe moderately sclerotized and developed; subapicodorsal lobe membranous, small, not reaching anterior edge, apex set on right side in dorsal view; right preapical edge deeply notched, basal edge of emargination produced into a sickle-shape projection; apex of apical lobe round apically, narrow and directed at right angle on the left side, apical villi regular in shape; subapicoventral lobe large, round, and interrupted in line with apical lobe. Variation. Measurements ( 117 ♂♂, 123 ♀♀). Length: male 7.5–11.5 mm(9.9 ± 1.0 mm), female 7.5–12.0 mm (10.0 ± 0.8 mm). Female allotype. Cephalic outline in dorsal view as in Fig. 76; vertex with a broadly arcuate and slightly sinuous medially transverse carina, dorsal edge slightly sinuous in frontal view (Fig. 97), lateral portion subangular and sloping down posteriorly; anterior pronotal tubercles well developed, external lateral edges nearly straight and slightly widening toward apex in dorsal view (Fig. 98), anterolateral surface obliquely concave, anterosuperior edge slightly arcuate in dorsal view, lateral portion of anterosuperior edge slightly upturned. Protibia short, with external teeth robust. Primary typedata. Holotype ♂( CMNC): [ BURKINA FASO: LORUM/ Forêt de Boulon, 270m/ 10°16’27”N004°27’15”O / 6.VII.2006, zone soudanienne / savane boisée, piège lumineux / F. & S. Génier, 2006- 05]; [COLÉOPTÈRES / DU / BURKINA FASO/ BF015489] barcode label; [ HOLOTYPE ♂/ Digitonthophagus/ falciger n.sp./ des. F. Génier, 2016] red card. MAP 7.Distribution of Digitonthophagus falciger.
Materialexamined( 292 ♂♂, 159 ♀♀), distribution (Map 7): BENIN: MONO, Agoué( 6°15'N, 1°41'E), vii– ix.1878, AbbéMénager— 2 ♀♀, 2 ♂♂( 4 paratypes) ( FGIC, MNHN); Agoué( 6°15'N, 1°41'E), 1879, AbbéMénager— 4 ♀♀, 18 ♂♂( 22 paratypes) ( FGIC, MNHN); BURKINA FASO: CENTRE, Ouagadougou( 12°22'N, 1°31'W), viii.1977, P. Ardoin— 1 ♂( paratype) ( FETC); Ouagadougou( 12°22'N, 1°31'W), 1977, J. Remy— 1 ♂( paratype) ( PMOC); CENTRE-NORD, 19 kmN Kaya, 314 m( 13°8'46.2''N, 0°52'42.1''W), 16.viii.2012, P. Moretto— 1 ♂( paratype) ( PMOC); CENTRE-OUEST, Forêt de Sorobouli[= Forêt Classée de Baporo], 270 m( 11°47'44''N, 2°53'25''W), 10–15.viii.2005, P. Moretto— 6 ♀♀, 3 ♂♂( 9 paratypes) ( PMOC); CENTRE-SUD, Réserve de Nazinga( 11°9'N, 1°36'W), 9–12.vi.2005, S. Rojkoff— 1 ♂( paratype) ( PCSR); EST, Kompienga( 11°4'26''N, 0°42'8''E), 3–24.ix.1988, F. Génier& M. Sanborne— 19 ♀♀, 47 ♂♂( 66 paratypes) ( CMNC); Kompienga, 20 kmS Pama( 11°4'37''N, 0°40'50''E), 1–16.vi.1988, Sanborne, Landry& Tou— 2 ♀♀( 2 paratypes) ( CMNC); Kompienga, 20 kmS Pama( 11°4'37''N, 0°40'50''E), 3–24.ix.1988, F. Génier& M. Sanborne— 2 ♀♀, 2 ♂♂( 4 paratypes) ( BDGC); HAUTS-BASSINS, Bobo-Dioulasso( 11°12'N, 4°18'W), 11.i.1975, H. Pollitzar— 1 ♂( paratype) ( BMNH); Bobo-Dioulasso( 11°12'N, 4°18'W), 14.xi.1976, R.H. Gooding— 3 ♂♂( 3 paratypes) ( CMNC); Matourkou( 11°5'N, 4°22'W), 15.vi.2005, S. Rojkoff— 1 ♂( paratype) ( PMOC); NORD, 8 kmSW Yako, Passoré, 320 m( 12°51'44''N, 2°18'35''W), 7.viii.2005, P. Moretto— 1 ♂( paratype) ( PMOC); SUD-OUEST, Bondigui, 324 m( 10°53'56.5''N, 3°34'15''W), v.2012, F. Rousset— 1 ♂( paratype) ( PMOC); Bondigui, 324 m( 10°53'56.5''N, 3°34'15''W), 20.viii.2012, P. Moretto— 2 ♀♀, 1 ♂( paratypes) ( PMOC); Gaoua( 10°19'N, 3°10'W), 1924, H. Labouret— 1 ♂( paratype) ( MNHN); COMOÉ, Forêt de Boulon(savane boisée), 270 m( 10°16'27''N, 4°27'15''W), 6.vii.2006, F. & S. Génier(2006-05)— 1 ♀, 1 ♂( holotype, allotype) ( CMNC); Forêt de Boulon(savane boisée), 270 m( 10°16'27''N, 4°27'15''W), 9.vii.2006, F. & S. Génier( 2006-17)— 2 ♂♂( 2 paratypes) ( FGIC); Forêt de Boulon(savane boisée), 270 m( 10°16'27''N, 4°27'15''W), 10.vii.2006, F. & S. Génier( 2006- 24)— 1 ♂( paratype) ( FGIC); KOMPIENGA, Kompienga, 160 m( 11°5'5''N, 0°42'42''E), 21.viii.2005, F. Génier( 2005-39)— 2 ♀♀, 5 ♂♂( 7 paratypes) ( FGIC); Pama, 230 m( 11°17'N, 0°42'59''E), 24.viii.2005, F. Génier( 2005- 44)— 1 ♂( paratype) ( FGIC); NAHOURI, Forêt de Nazinga, Kouzougou, 285 m( 11°9'17''N, 1°32'10''W), 21.vii.2006, F. & S. Génier( 2006-47)— 1 ♂( paratype) ( FGIC); Forêt de Nazinga, Kouzougou, 285 m( 11°9'17''N, 1°32'10''W), 25.vii.2006, F. & S. Génier( 2006-70)— 1 ♂( paratype) ( FGIC); Tiakané, 340 m( 11°11'3''N, 1°12'28''W), 18.viii.2005, F. Génier( 2005-23)— 1 ♀, 3 ♂♂( 4 paratypes) ( FGIC); Tiakané, 340 m( 11°11'3''N, 1°12'28''W), 20.viii.2005, F. Génier( 2005-34)— 1 ♂( paratype) ( FGIC); Tiakané, 340 m( 11°11'3''N, 1°12'28''W), 24.vii.2006, F. & S. Génier( 2006-64)— 1 ♂( paratype) ( FGIC); CAMEROON: EXTREME NORTH, Maroua( 10°35'N, 14°19'E), vii.1979, G. Onore— 1 ♀, 2 ♂♂( 3 paratypes) ( PHWC); Waza National Park( 11°15'N, 14°39'E), 18.viii.2006, C. Vanderbergh— 2 ♂♂( 2 paratypes) ( PMOC); NORTH, Garoua( 9°18'N, 13°24'E), x.1973, B. de Miré— 1 ♀, 2 ♂♂( 3 paratypes) ( MNHN); Garoua( 9°18'N, 13°24'E), [no date], [anonymous]— 1 ♀, 1 ♂( paratypes) ( OMOC); CHAD: HADJER-LAMIS, Sitje, rive sud Lac Tchad( 12°53'N, 14°52'E), 11.viii.2006, C. Vanderbergh— 1 ♀, 1 ♂( paratypes) ( PMOC); N’DJAMENA, Farcha( 12°7'N, 14°59'E), vi.1968, [anonymous]— 1 ♀, 2 ♂♂( 3 paratypes) ( MNHN); Farcha( 12°7'N, 14°59'E), 28.vi.1967, [anonymous]— 1 ♀, 1 ♂( paratypes) ( MNHN); Fort Lamy[= N’Djamena_] ( 12°7'N, 15°3'E), [no date], M. André— 1 ♀( paratype) ( PMOC); Fort Lamy[= N’Djamena_] ( 12°7'N, 15°3'E), 17.vii.1958, J. Mateu— 2 ♂♂( 2 paratypes) ( FETC); CÔTE D’IVOIRE: BAFING, Ranch de Dolla, Biémasso( 8°4'N, 7°33'W), 1–5.vi.2000, P. Moretto— 2 ♀♀, 14 ♂♂( 16 paratypes) ( PMOC); LACS, Bringakronear ( 6°25'53''N, 5°4'32''W), 12.vi.2003– 13.vi.2006, Newman et al.— 2 ♀♀, 3 ♂♂( 5 paratypes) ( BMNH); Bringakronear ( 6°25'53''N, 5°4'32''W), 13. vi.20 0 3, Newman et al.— 2 ♀♀, 4 ♂♂( 6 paratypes) ( BMNH); Bringakronear ( 6°25'53''N, 5°4'32''W), 17.vi.2003, Newman et al.— 1 ♂( paratype) ( BMNH); LAGUNES, 5 km Lamto( 6°13'N, 5°1'W), viii.1968, C. Girard— 2 ♀♀, 3 ♂♂( 5 paratypes) ( MNHN); Forêt du Banco( 5°23'N, 4°3'W), ii.1989, [anonymous]— 1 ♂( paratype) ( CEMT); NZI-COMOE, Dimbroko( 6°39'N, 4°42'W), [no date], [anonymous]— 1 ♂( paratype) ( IRSNB); SAVANES, Koko, Korhogo( 9°26'1''N, 4°43'9''W), 24–25.vi.2000, P. Moretto— 1 ♂( paratype) ( PMOC); VALLÉE DU BANDAMA, Bouaké( 7°41'N, 5°2'W), iii.1976, R. Couilloud— 2 ♀♀, 1 ♂( paratypes) ( PMOC); Bouaké( 7°41'N, 5°2'W), iv.1976, R. Couilloud— 2 ♀♀, 7 ♂♂( 9 paratypes) ( PMOC); Bouaké( 7°41'N, 5°2'W), 13.vii.1977, [anonymous]— 1 ♀( paratype) ( PMOC); GAMBIA: CENTRAL RIVER, Lower Saloum District, Ballangharr( 13°39'N, 15°25'W), 23.viii.1997, D.J. Mann— 7 ♀♀, 7 ♂♂( 14 paratypes) ( OUMNH); Naini District, Kuntaur Agricultural Station( 13°40'N, 14°53'W), 9.ix.1997, D.J. Mann— 1 ♂( paratype) ( OUMNH); LOWER RIVER, Jarra WestDistrict, JenoiDEC ( 13°28'51''N, 15°33'43''W), 29.viii.1997, Mann& Woodcock— 2 ♀♀, 2 ♂♂( 4 paratypes) ( OUMNH); UPPER RIVER, Bassa Government Rest Home, Mansanjang Kunda( 13°16'53''N, 14°12'13''W), 26.viii.1997, D.J. Mann& B. Woodcock— 1 ♂( paratype) ( OUMNH); WEST COAST, Kombo District, Yundum Agricultural Station( 13°21'43''N, 16°39'43''W), vii.1997, D.J. Mann— 1 ♂( paratype) ( OUMNH); Kombo District, Yundum Agricultural Station( 13°21'43''N, 16°39'43''W), 19.viii.1997, D.J. Mann& B. A. Woodcock— 2 ♂♂( 2 paratypes) ( OUMNH); GHANA: “ Danish Guinea”, [no date], P. E. Isert— 1 ♀, 1 ♂( paratypes) ( ZMUC); GUINEA: BOFFA, RioPongo ( 10°10'N, 14°2'W), 1883–1884, P. Lutz— 1 ♂( paratype) ( MNHN); GUINEA-BISSAU: BAFATÁ, Chime [= Xime], RioGéba ( 11°58'N, 14°56'W), 1906, G. Favarel— 2 ♀♀, 2 ♂♂( 4 paratypes) ( MNHN); TOMBALI, Quebo( 11°32'N, 14°46'W), 25.vi–17.vii.1995, A. Serrano— 1 ♂( paratype) ( CEMT); MAURITANIA: GUIDIMAKA, [unspecified locality] ( 15°22'N, 12°6'W), 1908, G. Audan— 1 ♀, 8 ♂♂( 9 paratypes) ( FGIC, MNHN); NIGER: NIAMEY, Gorou Kirey( 13°25'34''N, 2°8'32''E), 22.i.1976, [anonymous]— 1 ♂( paratype) ( PHWC); TILLABERI, Kolo( 12°55'22''N, 1°57'24''E), 15.iii.1972, D. Rougon— 1 ♂( paratype) ( PHWC); NIGERIA: KADUNA, Kaduna( 10°31'23''N, 7°26'25''E), 4.v.1962, D.C. Eidt— 1 ♂( paratype) ( CNC); Samaru( 11°10'N, 7°38'E), 18–25.v.1970, P.H. Ward— 1 ♀, 3 ♂♂( 4 paratypes) ( BMNH); Samaru( 11°10'N, 7°38'E), 15–22.vi.1970, P.H. Ward— 1 ♂( paratype) ( BMNH); KANO, [unspecified locality] ( 11°59'47''N, 8°31'0''E), v.1952, W.E.S. Merrett— 1 ♂( paratype) ( BMNH); [unspecified locality] (11°59'47''N, 8°31'0''E), xi.1951, W.E.S. Merrett—1 ♂ (paratype) (BMNH); [unspecified locality] (11°59'47''N, 8°31'0''E), v.1953, W.E.S. Merrett—1 ♂ (paratype) (BMNH); [unspecified locality] (11°59'47''N, 8°31'0''E), vi.1954, W.E.S. Merrett—1 ♂ (paratype) (BMNH); NIGER, Badeggi( 9°3'N, 6°9'E), 19.iii.1972, E.W. Classey— 1 ♂( paratype) ( BMNH); OYO, Ibadan( 7°22'N, 3°56'E), 2.iv.1957, J.L.Gregory— 1 ♀, 1 ♂( paratypes) ( BMNH); SENEGAL: DAKAR, Dakar( 14°46'N, 17°22'W), 1939, P. Lepesme, R. Paulian, & A. Villiers— 1 ♂( paratype) ( MNHN); Hann( 14°43'14''N, 17°26'12''W), 1939, P. Lepesme, R. Paulian, & A. Villiers— 15 ♀♀, 15 ♂♂( 30 paratypes) ( MNHN); Keur Ndiaye Lô( 14°45'25''N, 17°14'11''W), 18.viii.2013, I. de Dinechin— 1 ♂( paratype) (CMD); FATICK, Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 22.vi.2009, A. Coache— 1 ♀, 3 ♂♂( 4 paratypes) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 23.vi.2009, A. Coache— 2 ♀♀, 3 ♂♂( 5 paratypes) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 24.vi.2009, A. Coache— 3 ♂♂( 3 paratypes) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 16.x.2009, A. Coache— 1 ♂( paratype) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 28.xi.2009, A. Coache— 1 ♀( paratype) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 20.iv.2010, A. Coache— 1 ♂( paratype) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 15.vii.2010, A. Coache— 1 ♀, 1 ♂( paratypes) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 22.vii.2010, A. Coache— 1 ♂( paratype) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 13.viii.2010, A. Coache— 1 ♀, 1 ♂( paratypes) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 14.viii.2010, A. Coache— 2 ♀♀( 2 paratypes) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 15.viii.2010, A. Coache— 1 ♂( paratype) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 16.viii.2010, A. Coache— 1 ♂( paratype) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 17.viii.2010, A. Coache— 2 ♂♂( 2 paratypes) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 21.ix.2010, A. Coache— 1 ♀, 3 ♂♂( 4 paratypes) (CAC); Réserve de Fathala, Parcnational du delta du Saloum( 13°39'N, 16°27'W), 22.ix.2010, A. Coache— 1 ♀( paratype) (CAC); KAFFRINE, Tamba( 13°53'49''N, 15°15'33''W), 2.vii.1948, [anonymous]— 2 ♂♂( 2 paratypes) ( CMNC, CNC); KAOLAK, Missirah, 45 m( 13°59'26''N, 15°7'4''W), 16.vii.2007, F. Génier(2007-01)— 2 ♂♂( 2 paratypes) ( FGIC); Missirah, 45 m( 13°59'26''N, 15°7'4''W), 17.vii.2007, F. Génier(2007-03)— 1 ♂( paratype) ( FGIC); Nioro du Rip( 13°44'43''N, 15°46'23''W), 3.viii.2008, P. Moretto— 1 ♂( paratype) ( PMOC); Nioro du Rip( 13°44'43''N, 15°46'23''W), 10–11.viii.2009, P. Moretto— 1 ♀, 1 ♂( paratypes) ( PMOC); KÉDOUGOU, Dinndefelou( 12°22'43''N, 12°19'30''W), 13–14.vii.2008, P. Moretto— 1 ♂( paratype) ( PMOC); KOLDA, Forêt de Bakor, Mahon( 12°51'N, 14°49'W), 31.vii–2.viii.2008, P. Moretto— 1 ♂( paratype) ( PMOC); Mahon( 12°51'22''N, 14°48'42''W), 22–23.xi.2009, P. Moretto— 2 ♀♀, 3 ♂♂( 5 paratypes) ( PMOC); Sare Gnako, 31 m( 12°44'16''N, 15°9'56''W), 31.vii.2007, F. Génier& P. Moretto( 2007-49)— 2 ♂♂( 2 paratypes) ( FGIC); Sothoto, 23 m( 12°39'55''N, 15°24'3''W), 31.vii.2007, Génier& Moretto( 2007-47)— 2 ♂♂( 2 paratypes) ( FGIC); LOUGA, Linguère( 15°24'N, 15°7'W), ix.1967, A. Descarpentries, T.Leye, & A. Villiers— 10 ♀♀, 6 ♂♂( 16 paratypes) ( MNHN); Ndam-Dam( 15°36'22''N, 16°23'35''W), xi.1967, A. Descarpentries, T.Leye, & A. Villiers— 4 ♀♀, 1 ♂( paratypes) ( MNHN); SAINT-LOUIS, Parc Nationaldes Oiseaux du Djoudj( 16°24'N, 16°14'W), x.1996, [anonymous]— 1 ♀, 3 ♂♂( 4 paratypes) ( PSCW); Richard-Toll( 16°28'N, 15°41'W), xi.1967, A. Descarpentries, T.Leye & A. Villiers— 30 ♀♀, 23 ♂♂( 53 paratypes) ( MNHN); SEDHIOU, Sedhiou( 12°42'N, 15°34'W), [no date], [anonymous]— 1 ♂( paratype) ( MNHN); Sedhiou( 12°42'N, 15°34'W), 14.vii.1947, [anonymous]— 1 ♂( paratype) ( MNHN); TAMBACOUNDA, Boutou, 52 m( 13°48'16''N, 13°34'47''W), 20.vii.2007, F. Génier(2007-06)— 1 ♂( paratype) ( FGIC); Dienoun Diala( 13°12'47''N, 13°6'44''W), 29.vii.1995, P. Moretto— 8 ♀♀, 4 ♂♂( 12 paratypes) ( PMOC); ZIGUINCHOR, Boukitimgo, 18 m( 12°26'41''N, 16°35'53''W), 4.viii.2007, F. Génier( 2007-56)— 1 ♂( paratype) ( FGIC); Boukitimgo, 18 m( 12°26'41''N, 16°35'53''W), 4.viii.2007, P. Moretto& F. Génier( 2007-58)— 1 ♂( paratype) ( FGIC); SIERRA LEONE: NORTHERN, Koinadugu( 9°32'9''N, 11°22'6''W), 23.v.1963, MissionENS-IFAN aux MontsLoma— 1 ♀, 4 ♂♂( 5 paratypes) ( MNHN); UGANDA: Nebbi District, Pamora, Padyeresubcounty ( 2°24'25''N, 31°11'40''E), v.2006, Localcollectors— 1 ♀, 1 ♂( paratypes) ( PMOC).
Etymology. Falciger(sickle + bear) is a Latin adjective in apposition referring to the distinctive configuration of the left edge of the frontolateral peripheral sclerite of this species. Natural history. The few specimens with data were collected from savanna, disturbed areas, and pastures in soudanian zone and from degraded Guinean Forest. Attracted to mercury vapor and ultraviolet light, some specimens were collected using pitfall traps baited with human or western red colobus ( Procolobus badius) dung, some specimens retrieved from cow, donkey, elephant, rhinoceros, zebra, and zebu dung.
Remarks. The Ugandan record based on two specimens need confirmation.
1455385315
[151,1162,1329,1354]
Foret de Boulon
Burkina Faso
270
10.274166
Comoe
21
-4.4541664
32
33
1
Comoe
holotype
1455385302
2006-07-06
CMNC, LORUM
Foret de Boulon & F. & S. Genier & F. Genier
Burkina Faso
270
10.274166
Primary
Foret de Boulon
33
34
2
2
holotype
1455385309
[199,995,1436,1461]
Benin
Material
33
34
451
159
292
1455385289
1878-07
1878-09-31
1878-07
FGIC, MNHN
Abbe
Benin
6.25
Agoue
1305
1.6833334
33
34
8
2
2
MONO
1455385346
FGIC, MNHN
Abbe
Burkina Faso
6.25
Agoue
1305
1.6833334
33
34
44
4
18
1455385338
1977-08
FETC
Burkina Faso
12.366667
Ouagadougou
1293
-1.5166667
33
34
1
1
CENTRE
paratype
1455385355
PMOC
Burkina Faso
12.366667
Ouagadougou
1293
-1.5166667
33
34
1
1
CENTRE
paratype
1455385312
2012-08-16
PMOC
Burkina Faso
314
13.146167
Kaya
1
-0.8783611
33
34
1
1
CENTRE-NORD
paratype
1455385326
2005-08-10
2005-08-15
2005-08-10
PMOC
Foret de Sorobouli & Foret Classee de Baporo
Burkina Faso
270
11.795556
Foret Classee de Baporo
21
-2.8902776
Foret de Sorobouli
33
34
18
6
3
CENTRE-OUEST
paratype
1455385347
2005-06-09
2005-06-12
2005-06-09
PCSR
Reserve de Nazinga
Burkina Faso
11.15
Reserve de Nazinga
1296
-1.6
33
34
1
1
CENTRE-SUD
paratype
1455385322
1988-09-03
1988-09-24
1988-09-03
CMNC
F. Genier
Burkina Faso
11.073889
Kompienga
21
0.7022222
33
34
132
19
47
EST
paratype
1455385330
1988-06-01
1988-06-16
1988-06-01
CMNC
Sanborne & Landry
Burkina Faso
11.076944
Kompienga
21
0.6805556
33
34
4
2
EST
paratype
1455385358
1988-09-03
1988-09-24
1988-09-03
BDGC
F. Genier
Burkina Faso
11.076944
Kompienga
21
0.6805556
33
34
8
2
2
EST
paratype
1455385331
1975-01-11
BMNH
Burkina Faso
11.2
Bobo-Dioulasso
1295
-4.3
33
34
1
1
HAUTS-BASSINS
paratype
1455385336
1976-11-14
CMNC
Burkina Faso
11.2
Bobo-Dioulasso
1295
-4.3
33
34
6
3
HAUTS-BASSINS
paratype
1455385349
[267,1155,1904,1929]
2005-06-15
PMOC
Burkina Faso
11.083333
Matourkou
1296
-4.366667
33
34
1
1
HAUTS-BASSINS
paratype
1455385327
2005-08-07
PMOC
Burkina Faso
320
12.862223
Passore
21
-2.3097222
33
34
1
1
NORD
paratype
1455385318
2012-05
PMOC
Burkina Faso
324
10.899028
Bondigui
15
-3.5708332
33
34
1
1
SUD-OUEST
paratype
1455385316
2012-08-20
PMOC
Burkina Faso
324
10.899028
Bondigui
15
-3.5708332
33
34
3
2
1
SUD-OUEST
paratype
1455385344
MNHN
Burkina Faso
34
35
10.316667
Gaoua
1298
-3.1666667
33
34
1
1
SUD-OUEST
paratype
1455385337
2006-07-06
CMNC
Foret de Boulon & F. & S. Genier
Burkina Faso
270
10.274166
Foret de Boulon
21
-4.4541664
34
35
2
1
1
COMOE
holotype
1455385345
2006
2017
2006
FGIC
Foret de Boulon & F. & S. Genier
Burkina Faso
270
10.274166
Foret de Boulon
21
-4.4541664
34
35
4
2
COMOE
holotype
1455385343
2006
2024
2006
FGIC
Foret de Boulon & F. & S. Genier
Burkina Faso
270
10.274166
Foret de Boulon
21
-4.4541664
34
35
1
1
COMOE
paratype
1455385329
2005
2039
2005
FGIC
F. Genier
Burkina Faso
160
11.084722
Kompienga
21
0.71166664
34
35
14
2
5
COMOE
paratype
1455385350
2005
2044
2005
FGIC
F. Genier
Burkina Faso
230
11.283334
Pama
925
0.7163889
34
35
1
1
COMOE
paratype
1455385341
2006
2047
2006
FGIC
Foret de Nazinga & F. & S. Genier
Burkina Faso
285
11.154722
Kouzougou
21
-1.5361111
Foret de Nazinga
34
35
1
1
COMOE
paratype
1455385328
2006
2070
2006
FGIC
Foret de Nazinga & F. & S. Genier
Burkina Faso
285
11.154722
Kouzougou
21
-1.5361111
Foret de Nazinga
34
35
1
1
COMOE
paratype
1455385335
2005
2023
2005
FGIC
F. Genier
Burkina Faso
340
11.184167
Tiakane
21
-1.2077779
34
35
8
1
3
COMOE
paratype
1455385314
2005
2034
2005
FGIC
F. Genier
Burkina Faso
340
11.184167
Tiakane
21
-1.2077779
34
35
1
1
COMOE
paratype
1455385372
2006
2064
2006
FGIC
F. & S. Genier
Burkina Faso
340
11.184167
Tiakane
21
-1.2077779
34
35
1
1
COMOE
paratype
1455385363
1979-07
EXTREME, PHWC
Cameroon
10.583333
Maroua
1297
14.316667
34
35
6
1
2
NORTH
paratype
1455385380
2006-08-18
PMOC
Cameroon
11.25
Waza National Park
1295
14.65
34
35
4
2
NORTH
paratype
1455385334
1973-10
MNHN
Cameroon
9.3
Garoua
1300
13.4
34
35
6
1
2
NORTH
paratype
1455385352
OMOC
Chad
9.3
Garoua
1300
13.4
34
35
2
1
1
paratype
1455385371
2006-08-11
PMOC
Chad
12.883333
Sitje
1292
14.866667
34
35
2
1
1
HADJER-LAMIS
paratype
1455385348
1968-06
MNHN
Chad
12.116667
Farcha
1293
14.983334
34
35
6
1
2
HADJER-LAMIS
paratype
1455385353
1967-06-28
MNHN
Chad
12.116667
Farcha
1293
14.983334
34
35
2
1
1
HADJER-LAMIS
paratype
1455385382
[402,1432,799,824]
PMOC
Chad
12.116667
N'Djamena
1293
15.05
Fort Lamy
34
35
1
1
HADJER-LAMIS
paratype
1455385356
[151,1337,836,861]
1958-07-17
FETC
Chad
12.116667
N'Djamena
1293
15.05
Fort Lamy
34
35
4
2
HADJER-LAMIS
paratype
1455385360
2000-06-01
2000-06-05
2000-06-01
PMOC
Ranch de Dolla
Ivory Coast
8.066667
Biemasso
1302
-7.55
Ranch de Dolla
34
35
32
2
14
paratype
1455385375
[396,1070,908,933]
2003-06-12
Ivory Coast
6.431389
Bringakro
21
-5.0755553
34
35
1
LACS
paratype
1455385351
2006-06-13
BMNH
Ivory Coast
Newman
34
35
10
2
3
LACS
paratype
1455385342
vi. 2
BMNH
Newman
Ivory Coast
6.431389
Bringakro
21
-5.0755553
34
35
12
2
4
LACS
paratype
1455385340
2003-06-17
BMNH
Newman
Ivory Coast
6.431389
Bringakro
21
-5.0755553
34
35
1
1
LACS
paratype
1455385368
[270,1432,1015,1040]
1968-08
MNHN
Ivory Coast
6.2166667
Lamto
1305
-5.016667
34
35
10
2
3
LAGUNES
paratype
1455385365
[151,1125,1052,1077]
1989-02
CEMT
Ivory Coast
5.383333
Banco
1305
-4.05
34
35
1
1
Banco
paratype
1455385370
IRSNB
Ivory Coast
6.65
Dimbroko
1304
-4.7
34
35
1
1
SAVANES
paratype
1455385369
2000-06-24
2000-06-25
2000-06-24
PMOC
Ivory Coast
9.433611
Korhogo
21
-4.7191668
34
35
1
1
Koko
paratype
1455385357
1976-03
PMOC
Ivory Coast
7.6833334
Bouake
1303
-5.0333333
34
35
3
2
1
VALLEE DU BANDAMA
paratype
1455385374
1976-04
PMOC
Ivory Coast
7.6833334
Bouake
1303
-5.0333333
34
35
18
2
7
VALLEE DU BANDAMA
paratype
1455385364
1977-07-13
PMOC
Ivory Coast
7.6833334
Bouake
1303
-5.0333333
34
35
1
1
VALLEE DU BANDAMA
paratype
1455385354
[394,759,1231,1256]
Gambia
CENTRAL RIVER
34
35
1
CENTRAL RIVER
paratype
1455385362
1997-08-23
OUMNH
Gambia
13.65
Ballangharr
1290
-15.416667
34
35
28
7
7
Lower Saloum District
paratype
1455385339
1997-09-09
OUMNH
Gambia
13.666667
Kuntaur Agricultural Station
1290
-14.883333
34
35
1
1
Naini District
paratype
1455385396
[999,1197,1304,1328]
Gambia
LOWER RIVER
34
35
1
LOWER RIVER
paratype
1455385386
1997-08-29
OUMNH
Mann
Gambia
13.480833
Jenoi
21
-15.561945
34
35
8
2
2
Jarra West District
paratype
1455385403
1997-08-26
OUMNH
D. J. Mann
Gambia
13.281388
Mansanjang Kunda
21
-14.203611
Bassa Government Rest Home
34
35
1
1
UPPER RIVER
paratype
1455385359
[738,914,1412,1436]
Gambia
WEST COAST
34
35
1
WEST COAST
paratype
1455385377
1997-07
OUMNH
Gambia
13.361945
Yundum Agricultural Station
21
-16.661943
34
35
1
1
Kombo District
paratype
1455385397
1997-08-19
D. J. Mann
Gambia
13.361945
Yundum Agricultural Station
21
-16.661943
34
35
1
Kombo District
paratype
1455385373
OUMNH
Gambia
A
34
35
4
2
A
paratype
1455385379
[290,1203,1519,1544]
ZMUC
Ghana
Danish Guinea
34
35
2
1
1
paratype
1455385402
[151,1065,1556,1581]
1883
1884
1883
MNHN
Ghana
10.166667
Rio
1298
-14.033334
34
35
1
1
Rio
paratype
1455385378
Guinea-Bissau
BAFATA
34
35
1
BAFATA
paratype
1455385387
[372,1432,1591,1616]
MNHN
Guinea-Bissau
11.966666
Rio
1294
-14.933333
34
35
8
2
2
Rio
paratype
1455385400
[151,895,1628,1653]
1995-06-25
1995-07-17
1995-06-25
Guinea-Bissau
11.533334
Quebo
1295
-14.766666
34
35
1
TOMBALI
paratype
1455385376
[927,1432,1628,1652]
CEMT
Guinea-Bissau
A
34
35
1
1
A
paratype
1455385367
FGIC, MNHN
Mauritania
15.366667
GUIDIMAKA
1285
-12.1
34
35
18
1
8
GUIDIMAKA
paratype
1455385366
1976-01-22
PHWC
Mauritania
13.426111
Gorou Kirey
21
2.1422224
34
35
1
1
NIAMEY
paratype
1455385393
1972-03-15
PHWC
Mauritania
12.922778
Kolo
21
1.9566667
34
35
1
1
TILLABERI
paratype
1455385390
[266,529,1772,1797]
Nigeria
KADUNA
34
35
1
KADUNA
paratype
1455385394
[542,1432,1772,1797]
1962-05-04
CNC
Nigeria
10.523055
Kaduna
21
7.440278
34
35
1
1
Kaduna
paratype
1455385395
[151,1213,1807,1832]
1970-05-18
1970-05-25
1970-05-18
BMNH
Nigeria
11.166667
Samaru
1295
7.633333
34
35
8
1
3
Kaduna
paratype
1455385383
1970-06-15
1970-06-22
1970-06-15
BMNH
Nigeria
11.166667
Samaru
1295
7.633333
34
35
1
1
Kaduna
paratype
1455385398
1952-05
BMNH
W.D. Edmonds
Nigeria
11.996389
KANO
21
8.516666
34
35
1
1
KANO
paratype
1455385388
[391,1357,1988,2013]
1972-03-19
BMNH
Niger
9.05
Badeggi
1300
6.15
34
35
1
1
paratype
1455385381
1957-04-02
BMNH
J.-P. Lumaret
Niger
35
36
7.366667
Ibadan
1303
3.9333334
34
35
2
1
1
OYO
paratype
1455385384
[1101,1355,151,176]
Senegal
DAKAR
35
36
1
DAKAR
paratype
1455385361
MNHN
P. Lepesme & R. Paulian
Senegal
14.766666
Dakar
1286
-17.366667
35
36
1
1
Dakar
paratype
1455385419
MNHN
P. Lepesme & R. Paulian
Senegal
14.720555
Hann
21
-17.436666
35
36
60
15
15
A
paratype
1455385411
2013-08-18
Senegal
14.756945
Keur Ndiaye Lo
21
-17.23639
35
36
1
1
A
paratype
1455385430
2009-06-22
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
1
FATICK
paratype
1455385385
2009-06-23
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
8
1
3
A
paratype
1455385407
2009-06-24
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
10
2
3
A
paratype
1455385420
2009-10-16
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
6
3
A
paratype
1455385399
2009-11-28
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
1
1
A
paratype
1455385405
2010-04-20
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
1
1
A
paratype
1455385429
2010-07-15
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
1
1
A
paratype
1455385401
2010-07-22
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
2
1
1
A
paratype
1455385410
2010-08-13
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
1
1
A
paratype
1455385424
2010-08-14
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
2
1
1
A
paratype
1455385406
2010-08-15
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
4
2
A
paratype
1455385392
2010-08-16
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
1
1
A
paratype
1455385391
2010-08-17
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
1
1
A
paratype
1455385421
2010-09-21
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
4
2
A
paratype
1455385416
2010-09-22
Reserve de Fathala & du Saloum
Senegal
13.65
Parc
1290
-16.45
Reserve de Fathala
35
36
8
1
3
A
paratype
1455385422
[868,1278,871,896]
Senegal
A
35
36
1
1
A
paratype
1455385418
1948-07-02
CMNC, CNC
Senegal
13.896944
Tamba
21
-15.259167
35
36
4
2
KAFFRINE
paratype
1455385408
2007-07-16
FGIC
F. Genier
Senegal
45
13.990556
Missirah
21
-15.117778
35
36
4
2
KAFFRINE
paratype
1455385425
2007-07-17
FGIC
F. Genier
Senegal
45
13.990556
Missirah
21
-15.117778
35
36
1
1
KAFFRINE
paratype
1455385414
2008-08-03
PMOC
Nioro du Rip
Senegal
13.745278
Nioro du Rip
21
-15.773055
35
36
1
1
KAFFRINE
paratype
1455385404
2009-08-10
2009-08-11
2009-08-10
PMOC
Nioro du Rip
Senegal
13.745278
Nioro du Rip
21
-15.773055
35
36
2
1
1
KAFFRINE
paratype
1455385409
2008-07-13
2008-07-14
2008-07-13
PMOC
Senegal
12.378612
Dinndefelou
21
-12.325
35
36
1
1
KEDOUGOU
paratype
1455385389
2008-07-31
2008-08-02
2008-07-31
PMOC
Foret de Bakor
Senegal
12.85
Mahon
1292
-14.816667
Foret de Bakor
35
36
1
1
KOLDA
paratype
1455385440
2009-11-22
2009-11-23
2009-11-22
PMOC
Senegal
12.856112
Mahon
21
-14.8116665
35
36
10
2
3
KOLDA
paratype
1455385433
2007
2049
2007
FGIC
F. Genier & P. Moretto
Senegal
31
12.737778
Sare Gnako
21
-15.165555
35
36
4
2
KOLDA
paratype
1455385455
2007
2047
2007
FGIC
Genier & Moretto
Senegal
23
12.6652775
Sothoto
21
-15.400833
35
36
4
2
KOLDA
paratype
1455385413
1967-09
Senegal
15.4
Linguere
1285
-15.116667
35
36
1
LOUGA
paratype
1455385436
1967-11
MNHN
Descarpentries, T.
Senegal
15.606112
Ndam-Dam
21
-16.393055
35
36
32
10
6
A
paratype
1455385449
MNHN
Descarpentries, T.
Senegal
A
35
36
5
4
1
A
paratype
1455385423
1996-10
PSCW
Oiseaux du Djoudj
Senegal
16.4
Oiseaux du Djoudj
1282
-16.233334
Parc National
35
36
8
1
3
SAINT-LOUIS
paratype
1455385428
1967-11
MNHN
Descarpentries, T.
Senegal
16.466667
Richard-Toll
1282
-15.683333
35
36
106
30
23
A
paratype
1455385453
[853,979,1412,1436]
Senegal
SEDHIOU
35
36
1
SEDHIOU
paratype
1455385426
MNHN
Senegal
12.7
Sedhiou
1292
-15.566667
35
36
1
1
Sedhiou
paratype
1455385434
1947-07-14
MNHN
Senegal
12.7
Sedhiou
1292
-15.566667
35
36
1
1
Sedhiou
paratype
1455385450
2007-07-20
FGIC
F. Genier
Senegal
52
13.804444
Boutou
21
-13.579722
35
36
1
1
TAMBACOUNDA
paratype
1455385435
1995-07-29
PMOC
Senegal
13.213056
Dienoun Diala
21
-13.112223
35
36
24
8
4
TAMBACOUNDA
paratype
1455385417
2007
2056
2007
FGIC
F. Genier
Senegal
18
12.444722
Boukitimgo
21
-16.598057
35
36
1
1
paratype
1455385415
2007
2058
2007
FGIC
P. Moretto & F. Genier
Senegal
18
12.444722
Boukitimgo
21
-16.598057
35
36
1
1
paratype
1455385445
1963-05-23
MNHN
Mission & Monts
Sierra Leone
9.535833
Koinadugu
21
-11.368334
35
36
10
1
4
NORTHERN
paratype
1455385439
2006-05
PMOC
Local
Uganda
2.4069445
Padyere
21
31.194443
Pamora
35
36
2
1
1
Nebbi District
paratype