Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)
Génier, François
Moretto, Philippe
Zootaxa
2017
4248
1
1
110
363WG
[151,475,1519,1545]
Insecta
Scarabaeidae
Digitonthophagus
Animalia
Coleoptera
14
15
Arthropoda
species
bonasus
Diagnosis. Male cephalic horns long, obliquely divergent in frontal view ( Figs. 24, 26, 28); females with horns on vertex ( Figs. 71–73).