Harpactocrates lycaoniae Harpactocrates lycaoniae Harpactocrates lycaoniae The genus Hygrocrates Deeleman-Reinhold, 1988 (Araneae, Dysderidae) in Turkey Kunt, Kadir Bogac Yagmur, Ersen Aydin Oezkuetuek, Recep Sulhi Kaya, Rahsen S. ZooKeys 2011 85 1 16 Brignoli, 1978 Brignoli 1978 Arachnida Dysderidae Hygrocrates CoL Animalia Hygrocrates lycaoniae Araneae 6 7 Arthropoda species lycaoniae   Harpactocrates lycaoniae: Brignoli, 1978: 463, f. 2, 8 (D ♂).  Harpactocrates lycaoniae.: Deeleman-Reinhold & Deeleman, 1988: 240, f. 13, 22, 320-325 (T ♂ from Harpactocrates, D ♀).  Harpactocrates lycaoniae.: Dunin, 1992: 41, f. 3 (♂ ♀).  Material Examined: 1 ♂, 1 ♀ (SMF) (abdomen heavily damaged during dissection) Burdur Province, YesilovaDistrict, side of Salda Lake [37°30'32.78"N, 29°41'56.66"E], 14.VII.2010, leg. E.A. Yagmur& M. Elverici.  Diagnosis:  Hygrocrates lycaoniaecan be distinguished from Hygrocrates caucasicusby the pyriform shape of the bulbus (bulbus smooth and cylindrical in Hygrocrates caucasicus) (see Dunin 1992); and from Hygrocrates georgicusby having apically oval-shaped spermathecae (see Mcheidze 1972).  Note:  Hygrocrates caucasicuswas originally described on the basis of two males by Dunin (1992). The females of this species have not been collected yet, but the bulbal structures of the male palp were well illustrated by Dunin (1992). However, the information of Hygrocrates georgicusis still insufficient. The following information was given by Dunin (1992: p. 60) inhis review of the Caucasian Dysderidae(our translation!): "H. georgicus was described on the basis of the female holotype from Georgia: the vicinity of Tbilisi, Kodzhori, Udzho. The holotype was lost. The species was tentatively placed in Hygrocates by Deeleman-Reinhold and Deeleman (1988). To confirm their placement addition material is required. This species is absent from my collection." During the preparation of this paper, as a result of our correspondence with the Tbilisi Janashia Museum (Georgia) which retains the spider collection of Tamara Mcheidze, it is obvious that the holotype of this species is lost (S. Otto pers. comm.). Thus, we could not examine it, but on the basis of the original illustrations of the vulva by Mcheidze (1972), Hygrocrates georgicuscan be distinguished from the Turkish members of the genus by the linear distalmost part of spermathecae.   Measurements(♂ / ♀): AL 3.36 /?; CL 2.40 / 3.00; CWmax 2.20 / 2.44; CWmin 1.52 / 1.76; AMEd 0.14 / 0.16; PLEd 0.13 / 0.15; PMEd 0.11 / 0.12; ChF 0.77 / 0.79; ChG 0.45 / 0.50; ChL 1.30 / 1.46. Leg measurements are given in Table 3.  Description: General features of the body of Hygrocrates lycaoniaeclosely resemble the new species (Figs 22-29), Hygrocrates deelemanussp. n., but the two are easily differentiated by their different body sizes and by structures of the male and female genitalia (Figs 30-33, 35-38). The males of the two species are easily distinguished in ventral view (90° angle) by the terminal part of the bulbus having the following characteristics: 1. Embolic base is pear-shaped in the two species, but the tip of the embolic base located at 12 o'clockin Hygrocrates deelemanussp. n. (Fig. 37) and at 10 o'clockin Hygrocrates lycaoniae(Fig. 38). 2. Apophysisa and Apophysisb are short and blunt in Hygrocrates deelemanussp. n. (Fig. 37), but long in Hygrocrates lycaoniae(Fig. 38).  3. Apophysisb originates near the tip of the embolic base in Hygrocrates deelemanussp. n. (Fig. 37), but originates from the central part of the tip of the embolic base in Hygrocrates lycaoniae(Fig. 38). 4.Embolus is curved between Apophysisa and Apophysisb in Hygrocrates deelemanussp. n. (Fig. 37), but is raised from the embolic base and separated from Apophysisa and Apophysisb in Hygrocrates lycaoniae(Fig. 38). The females of the two species are easily distinguished by the form of the proximalmost part of the spermathecae which is oval in Hygrocrates deelemanussp. n. (Figs 20-21) and circular in Hygrocrates lycaoniae(Fig. 34). Details of leg spination are given in Table 4.  Figures 22-29. Hygrocrates lycaoniae22 (♂), 26 (♀) carapace, dorsal view 23 (♂), 27 (♀) ditto, anterior view 24 (♂), 28 (♀) ditto, lateral view 25 (♂), 29 (♀) ditto, ventral view. Scale lines: (22, 24, 25, 26, 28, 29) 0.5 mm; (23, 27) 0.25 mm  Figures 30-33. Male palp of Hygrocrates lycaoniae. Abbreviations: Apa apophysisa Apb apophysisb E embolus. Scale line: 0.1 mm.  Figure 34. Vulva of Hygrocrates lycaoniae, dorsal view. Abbreviations: Ad anterior diverticulum Pd posterior diverticulum Sp spermatheca dps distalmost part of spermatheca pps proximalmost part of spermatheca da dorsal arch cv central valve tb transverse bar ps membranous sac tlm twisted lateral membranes. Scale line: 0.1 mm  Table 2. Leg spination of Hygrocrates deelemanussp. n.    ♂ (Holotype) Leg I Leg II Leg III Leg IV  Table 3. Leg measurements of Hygrocrates lycaoniae    (♂ / ♀) Fe Pa Ti Mt Ta  Figures 35-38. Comparison of the palps of the two species. 35, 37 Hygrocrates deelemanussp. n. 36, 38 Hygrocrates lycaoniae. Abbreviations:Apa apophysisa Apb apophysisb E embolus.