New Species Of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) From Mexico, Guatemala, Colombia, And Brazil Ratcliffe, Brett C. The Coleopterists Bulletin 2001 2001-12-01 55 4 433 443 Endrodi 1978: 91 Endrodi 91 1978 [246,703,340,361] Insecta Dynastidae Hemiphileurus Animalia Coleoptera 5 438 Arthropoda species rugulosus  To help alleviate this confusion, I am providing new illustrations (Figs. 11– 12) of the male genitalia of  H. rugulosusspecimens collected at Rancho Grande (near Maracay), Venezuelaas well as Endrödi’s original drawing (Fig. 13).   Hemiphileurus quadridentatusRatcliffe,  new species( Figs. 14–15)   Type Material.  Holotypelabeled ‘‘ GUATEMALA: Izabal, SE Moralesnr. Negro Norte,  IV­1997,;  1,000 m, 158229670N 888419680W, col. José Monzon.’’  Allotypewith same data. Six male and three female paratypeswith same data.  Holotypeand allotypedeposited at the Universityof Nebraska State Museum( Lincoln, NE).  Paratypesdeposited in the collections of the University of Nebraska StateMuseum, U.S.National Museum ( Washington, D.C. but currently at the University of Nebraskafor off­site enhancement), Universidad del Valle(Guatemala City, Guatemala), and the B.C. Ratcliffecollection.   Holotype. Male. Length 21.1 mm; width 9.7. Color black. Head: Surface of frons and base of clypeus coarsely rugose, apex of clypeus with sparse, moderately­sized punctures. Frons near its apex with 2 low, rounded tubercles; center of frons with large, moderately deep, oval fovea. Clypeus triangular, apex acute and strongly reflexed, carinae from apex to each tubercle absent. Interocular width equals 3.1 transverse eye diameters. Antenna with 10 segments, club subequal in length to segments 2–7. Mandibles arcuate on external edge, apices acute. Pronotum: Surface with disc either side of middle moderately punctate, punctures mostly large and weakly ocellate; punctures along anterior margin a little smaller, becoming rugopunctate, punctures on lateral margins moderate in size. Median, longitudinal furrow deep, with large and dense punctures, furrow extending from near base to near apex. A faint (nearly obsolete) tubercle present just behind apex either side of median furrow. All margins completely beaded. Elytra: Surface with 6 distinct, punctate striae between suture and humeral umbone; punctures moderate to large, oval to elongate, strongly ocellate, separated from one another in each row by about 1 puncture length. Intervals convex, impunctate.  Pygidium: Surfacemoderately densely punctate; punctures moderate to mostly large, ocellate, setigerous; setae moderate to mostly long, reddish brown. In lateral view, surface regularly convex. Legs: Foretibia quadridentate, teeth subequally spaced from one another. Median transverse carina on meso­ and metatibiae terminating with strong, acute spine. Apex of posterior tibia with strong, acute spine on upper angle and with 5 short spinules below spine. Apex of first tarsomere on posterior tarsus with strong, acute spine dorsally. Venter: Prosternal process broadly rounded, short, apex truncate and weakly concave. Last sternite rugopunctate in narrow band along base, elsewhere virtually impunctate. Parameres: Figures 14–15.   Allotype. Female. Length 21.9 mm; width 9.4 mm. As holotypeexcept in the following respects: Head: Clypeus with a small, weak carina extending from apex to base of   Figs. 14–15.Parameres of  H. quadridentatusRatcliffe, n. sp.  Figs. 16–17.Parameres of  H. warneriRatcliffe, n. sp.  Fig. 18.Prosternal process, lateral oblique view, of  H. warneriRatcliffe, n. sp.  Figs. 19–20.Parameres of  H. illatus(LeConte). each frontal tubercle. Elytra: Punctures of striae mostly round to oval, only a few elongate. Venter: Last sternite with basal 1/3 rugopunctate, with sparse micropunctures elsewhere.   Variation.Males (5). Length 19.5–21.8 mm; width 8.6–9.3 mm. The male paratypesdo not differ appreciably from the holotype. One specimen has fewer punctures on the pronotum and less setae on the pygidium. Females (3). Length 18.5–23.3 mm; width 8.5–10.3 mm. The female paratypesdo not differ from the allotypeexcept except for the presence of faint clypeal carinae, and the pygidiumis weakly convex with smaller punctures in one specimen.   Etymology.This species is descriptively named for the four teeth present on the foretibia.   Remarks.  Hemiphileurus quadridentatuswill key only to couplets 45/46, the last in Endrödi’s (1985) key. Here one finds the only other continental species with quadridentate foretibiae:  H. microps(Burmeister)and  H. illatus(LeConte), both from Mesoamerica. A third continental species with quadridentate foretibiae,  H. warneri,is described in this paper. The diagnostic male parameres must be examined for reliable identification because of overlapping external character states between the four species. The only other species with quadridentate foretibiae, out of 43 (currently) in the genus, are  H. dispar(Kolbe)and  H. parvusDupuis and Dechambre, from Hispaniola and Cubarespectively. The specimens of  H. quadridentatuswere taken at lights along with several specimens of  H. microps; nothing is known of their biology. Although the typelocality near Negro Norte is only 1,000 min elevation, it is a cloud forest because of the high amount of moisture received from the Caribbean (. 4,000 mmof rain/year). According to J. Monzon (pers. comm., September 2000), this forest is the northernmost limit for many Central and South American Cerambycidaeas well as the metallic species of Central American  Plusiotis( Scarabaeidae: Rutelinae) and other beetles. This is one of the most biologically rich forests in Guatemalaand should be considered in any conservation programs there. Venezuela Venezuela 5 438 43 [428,816,409,430] Colombia Colombia 5 438 1 1997-04 SE Morales & Jose Monzon. Guatemala 1000 Negro Norte 5 438 1 Izabal holotype [246,1075,881,902] 1997-04 SE Morales & Jose Monzon. Guatemala 1000 Negro Norte 5 438 2 1 Izabal allotype Lincoln, NE Guatemala Museum University 5 438 2 Nebraska holotype B. C. Ratcliffe United States of America Universidad del Valle 5 438 1 Nebraska paratype