Laeocathaica polytyla Laeocathaica (Laeocathaica) polytyla Land snail diversity in central China: revision of Laeocathaica Moellendorff, 1899 (Gastropoda, Camaenidae), with descriptions of seven new species Wu, Min Shen, Wang Chen, Zhong-Guang ZooKeys 2023 2023-03-20 1154 49 147 7RSP9 2821039 59689614-5308-5EFF-BB49-6202AE40C409 Moellendorff, 1899 Moellendorff 1899 Gastropoda Camaenidae Laeocathaica CoL Animalia Laeocathaica polytyla Stylommatophora 0 49 Mollusca species polytyla   Figs 2A , 3 , 24A, B , 26 , 43A, B , 45K, L , 51   Laeocathaica polytyla Moellendorff, 1899: 98, pl. 6, fig. 7; - Wiegmann 1900: 118, pl. 3, figs 105-107; - Gude 1902: 6; - Yen 1939: 150, pl. 15, fig. 41; - Schileyko 2004: 1686, fig. 2174B-D; - Chen and Zhang 2004: 331, fig. 322; -  Pall-Gergelyet al. 2022: 68, fig. 20A-C.  Laeocathaica (Laeocathaica) polytyla- Zilch 1968: 174; - Richardson 1983: 78.  Museum material. SMF 9098, lectotype; Nanping, Sung-pan. SMF 9099, paratypes; six fms; Nanping, Sung-pan. Other SMF material see in Yen 1939(150).  New material.  HBUMM00506, 13 fma, 1 dissected; Anchanghexiang[安昌河乡], Wenxian, Gansu Province,  1200 m, near point ( 33.066003°N, 104.460018°E); 1998-V-19, coll. Chen, D.-N.and Zhang, G.-Q. HBUMM00513, a subadult; Shuanghexiang[双河乡], Jiuzhaigouxian, Sichuan Province,  1100 m, near ( 33.179715°N, 104.264968°E); 1998-V-18, coll. Chen, D.-N.and Zhang, G.-Q. HBUMM00579, many fms; Jiuzhaigouxian, Sichuan Province; 1998-V-18, coll. Chen D.and Zhang, G.-Q. HBUMM05419, 1 fms; hill foot of Yuxushan, Wenxian, Gansu Province; slate and shrubs, dry; 2006-IX-27, coll. Wu, M., Liu, J.-M., Zheng, W.and Gao, L.-H. HBUMM06490, many fms; Yuxushan, Wenxian, Gansu Province,  1070 ma.s.l., 32.959861°N, 104.678222°E, shrubs and slate, 2011-VIII-07, coll. Wu, M., Xu, Q.and Budha, P. HBUMM06523, many fms and juvs; town of Wenxian, Gansu Province,  1024 ma.s.l., 32.941111°N, 104.668889°E; 2011-VIII-08, coll. Wu, M., Xu, Q.and Budha, P. HBUMM08454, 23 fms, east of the town of Jiuzhaigouxian, Sichuan Province, 2020-VIII, coll. Chen, Z.-G. CZG202008-w6, 1 subadult; Shangdezhen, Wenxian, Gansu Province, near point ( 32.907414°N, 104.76994°E); 2020-VIII, coll. Chen, Z.-G. CZG202008-w11, 2 fms, near point ( 32.94936°N, 104.693006°E), town of Wenxian, Gansu Province, 2020-VIII, coll. Chen, Z.-G. HBUMM05437, many fma; 1 fma dissected; Yuxushan, Wenxian, Gansu Province, 32.957259°N, 104.689152°E, shrubs and slate, 2006-IX-27, coll. Wu, M., Liu, J.-M., Zheng, W.and Gao, L.-H.; DNA voucher HBUMM05411. HBUMM06753, 3 fma, 7 fms; Easternbank of Baishuijiang, Lihuacun, Jiuzhaigouxian, Sichuan Province,  1425 m, 33.267222°N, 104.234722°E; 2011-VI-14, coll. Wu, M., Xu, Q.and Buhda, P.   Figure 24. A, B  Laeocathaica polytyla Moellendorff, 1899 ASMF 9098, lectotype BHBUMM05437-spec.1 C, D  L. potanini Moellendorff, 1899 CSMF 9082, paratype DHBUMM00633-spec.1.  Distribution. Gansu: Wenxian. Sichuan: Jiuzhaigouxian (type locality).  Additional information of shell. Tiny elongate granules on protoconch, which can be only observed in juvenile shells, are densely and radially arranged. The granules on protoconch are absent in adult shell because of erosion or weathering. On teleoconch the growth lines are indistinct. Spiral grooves are absent throughout.   Figure 25.  Laeocathaica parapolytylaWu, sp. nov. A-Eparatypes HBUMM06640-spec.2-6 Fholotype, HBUMM06640-spec.1 G-Iparatypes HBUMM06640-spec.7-9. Both A, Hshow four views B-G, Ishow apical view only.  General anatomy. Eversible head wart prominent. Jaw arcuate, with three projecting ribs.  Anatomy of genital organs. Penial sheath covering ~ 1/4 of penis. Penis proximately tubular and distally expanded. Inside penis, two pilasters fusing into one Y-shaped fork at proximal 1/5, two more pilasters of similar thickness also present. Fine pilasters on distal end of penis merging into ~ 8 thick folds. Vas deferens narrow throughout. Vagina between atrium and dart sac not elongated. Vagina between dart sac and insertion of bursa copulatrix duct ~ 1/2 length of dart sac. Dart sac ~ 1/2 length of penis. Love dart ~ 1 mm long, apically 2-bladed. Accessory sac small but externally distinguishable, internally solid, inserting into dart sac at distal 1/3, opening to dart sac chamber. Mucous glands four or five, each singly tubular or simply branched. Proximal accessory sac absent. Bursa copulatrix duct equally thick.   Figure 26.Genital anatomy of  Laeocathaica polytyla Moellendorff, 1899, HBUMM05437-spec.1 Ageneral view Bcross-section of dart sac at the position arrowed in ( A) C, Dleft and ventral side of dart sac apparatus. Abbreviations: AS - accessory sac; At - atrium; BCD - bursa copulatrix duct; DS - dart sac; DSC - dart sac chamber; DtC - a chamber containing love dart; Ep - epiphallus; FO - free oviduct; MG - mucous glands; P - penis; PR - penial retractor muscle; PS - penial sheath; Va - vagina; VD - vas deferens.  Remarks. See  Laeocathaica parapolytylaWu, sp. nov. 1998-05-19 Chen, D. - N., Zhang, G. - Q. Gansu Province 1200 33.066 Anchanghexiang 1 104.460014 Wenxian 1 1998-05-18 Chen, D. - N., Zhang, G. - Q. Sichuan Province 1100 33.179714 Shuanghexiang 1 104.26497 Jiuzhaigouxian 1 1998-05-18 Chen D., Zhang, G. - Q. Jiuzhaigouxian Sichuan Province 1 2006-09-27 Wu, M., Liu, J. - M., Zheng, W., Gao, L. - H. Gansu Province Yuxushan Wenxian 1 2011-08-07 Wu, M., Xu, Q., Budha, P. Gansu Province 1070 32.95986 Yuxushan 1 104.67822 Wenxian 1 2011-08-08 Wu, M., Xu, Q., Budha, P. 1024 32.941113 Wenxian 1 104.66889 Gansu Province 1 2020-08 Chen, Z. - G. Jiuzhaigouxian Sichuan Province 1 2020-08 Chen, Z. - G. Gansu Province 32.907413 Shangdezhen 1 104.76994 Wenxian 1 1 2020-08 Wenxian, Chen, Z. - G. 32.94936 Gansu Province 1 104.69301 1 2006-09-27 Wu, M., Liu, J. - M., Zheng, W., Gao, L. - H. Gansu Province 32.95726 Yuxushan 1 104.689156 Wenxian 1 2011-06-14 Wu, M., Xu, Q., Buhda, P. Sichuan Province 1425 33.267223 Eastern 1 104.23472 Jiuzhaigouxian 1