Bryaxis valdiviensis Reitter, 1883: 50 valdiviensis Blanchard, 1851 Reitter, 1885: 326 Achilia clavata Raffray, 1904: 137 Jeannel, 1962: 429 Achilia blanchardi Raffray, 1904: 138 Bryaxis valdiviensis Reitter, 1883 Jeannel, 1962: 426 A revision of the Chilean Brachyglutini - Part 7. Revision of Achilia Reitter, 1890: A. cosmoptera species group (Coleoptera: Staphylinidae: Pselaphinae) Kurbatov, Sergey A. Cuccodoro, Giulio Sabella, Giorgio Revue suisse de Zoologie 2021 2021-04-23 128 1 135 156 9776 Raffray, 1904 Raffray 1904 [950,1317,419,443] Insecta Staphylinidae Achilia Animalia Coleoptera 5 139 Arthropoda species blanchardi   Figs 3, 17, 27, 38, 40, 42, 63      Bryaxis valdiviensis Reitter, 1883: 50, pl. 1 fig. 8 (head and antenna) (nec  valdiviensis Blanchard, 1851). –  Reitter, 1885: 326; 330, pl. 2 fig. 14 (head and antenna).     Achilia clavata Raffray, 1904: 137, fig. 53 (head and antenna) (synonymized by  Jeannel, 1962: 429).     Achilia blanchardi Raffray, 1904: 138(new name for  Bryaxis valdiviensis Reitter, 1883). –  Jeannel, 1962: 426, 430, figs 195 (head and antenna of male), 196 (head and antenna of female), 197 (aedeagus).   Type material ( 12 ex.):  MNHN; 1 ♀( Holotypeof  A. clavata); labels verbatim “Museum de Paris; 1917; coll. Raffray / Chili/ Type /  clavata; det. A. Raffray/  clavata(handwritten by Jeannel)”. – SOUTHERN CHILE: Región Los Ríos: Valdivia Prov.: MNHN; 1 ♂( lectotypeof  A. blanchardihere designated); labels verbatim “Lectotype / Type / 1880; Chili; Valdivia; leg. Kindermann/  Bryaxis;  valdiviensis; m. Valdivia / Museum de Paris; 1917; coll. Raffray/  A. blanchardi; A. Raffraydet.”. – MNHN; 1 ♂and 9 ♀( Paralectotypesof  A. blanchardihere designated); labels verbatim “Paralectotype / Chili/ Museum de Paris; 1917; coll. Raffray/  A. blanchardi; A. Raffraydet.”.  Additional material examined ( 5 ex.):  FMNH; 1 ♀; Field Mus. Nat. Hist.; Orlando Park; Pselaphidae Colln.; F. C. Fletcher Collection. – MNHN; 1 ♀; Chile; P. Germain. – HNHM; 1 ♂; “ Chili”. – MNHN; 2 ♀; “ Chili”; Gay; 1849.   Description:Body 1.35-1.40 mmlong, dark brown with reddish elytra; antennae and legs reddish; palpi yellowish.  Male: Head as in Figs 38, 40 & 42, very wide, surface with some very scattered punctures; occipital region raised, frons flattened and separated by large transverse sulcus from large frontal lobe; anterior margin of frontal sulcus slightly raised and pointed at middle. Antennae ( Fig. 17) with scape and pedicel longer than wide; antennomere III distinctly longer than wide; antennomere IV about as long as wide; antennomeres V-VII slightly longer than wide; antennomere VIII as long as wide; antennomere IX wider than VIII and wider than long; antennomere X distinctly wider than long, shorter and wider than IX; antennomere XI elongate, longer than VII-X combined. Metaventrite with deep and wide transverse sulcus on apical twothirds, sulcus with deflected and posteriorly prominent margins forming sort of tubercles. First abdominal sternite slightly raised and flattened at middle; remaining abdominal sternites slightly flattened at middle. Protibiae with distal half slightly swollen; mesotibiae ( Fig. 27) with distal half swollen and densely pubescent. Aedeagus ( Fig. 3) 0.23 mmlong, with dorsal plate subrectangular; dorsal longitudinal struts divergent. Parameres relatively wide with very large and long recurved seta on welldeveloped outer lobe, this seta bifid and very swollen on basal third; apical portion of parameres slightly recurved anteriorly and prolonged laterally at tip; apex of parameres bearing one long ventral medial seta associated with wider and shorter seta. Copulatory pieces consisting of two subequal short sclerites slightly enlarged apically, recurved and more robust basally.  Female: Similar to male except head not modified; antennae shorter with antennomere XI shorter; metaventrite, abdominal sternites, and legs unmodified.   Collecting data:The data in our possession do not allow us to make statements in this regard.   Distribution:  Achilia blanchardiis only known from Valdivia ( Región Los Ríos: Valdivia Prov.) ( Fig. 63: green diamonds).   Comments: Raffray (1904: 138)proposed the new name  Achilia blanchardifor  Bryaxis valdiviensis Reitter, 1883, described on an unspecified number of specimens from Valdivia, because this name was preoccupied by  Pselaphus valdiviensis Blanchard, 1851[now  Achilia valdiviensis( Blanchard, 1851)] (see Sabella et al., 2020: 140, 144-145).  Jeannel (1962: 430)listed  Achilia clavata Raffray, 1904as a junior synonym of  A. blanchardi, but without justifying this action. We have examined the holotypeof  A. clavata, which is a female resembling in all features those of  A. blanchardi, and therefore we confirm the synonymy proposed by Jeannel.  Jeannel (1962: 430)stated that the type of  Achilia blanchardi Raffray, 1904was housed in the MNHN collections. In the MNHN we found in Raffray’s collection a series of 11 specimensidentified by Raffray as  Achilia blanchardi; the first specimen of the series is a male labeled “Type / 1880; Chili; Valdivia; leg. Kindermann/  Bryaxis;  valdiviensis; m. Valdivia / Museum de Paris; 1917; coll. Raffray /  A. blanchardi; A. Raffray det.” which we designate here as the lectotypeof  A. blanchardi Raffray, 1904( Blanchard, 1851), and the remaining 10 specimens( 1 maleand 9 females) labeled “ Chili” without further label data are designated as paralectotypes. Within the  A. cosmopteragroup the males of  A. blanchardiare easily distinguished from other species by the shape of the head ( Figs 38, 40, 42), antennae ( Fig. 17) and aedeagus ( Fig. 3). The females of this species are very similar to those of  A. elfridae, from which they can be distinguished by their wider head with a wider frontal sulcus, and their slightly longer antennomere XI ( 0.16 mmfor  A. blanchardi vs 0.14-0.15 mmfor  A. elfridae). 3397993412 MNHN Raffray Chile A. Raffray 5 139 1 1 holotype 3397993368 MNHN Raffray Chile Type A. Raffray Valdivia Prov. Valdivia Prov. 5 139 1 1 Region Los Rios lectotype 3397993385 MNHN Raffray Chile A. Raffray 5 139 10 9 1 Los Rios paralectotype 3397993356 FMNH Field Mus. Nat. Hist. Orlando Park 5 139 1 1 3397993464 [944,1281,1169,1191] MNHN P. Germain Chile 5 139 1 1 3397993450 HNHM Chile 5 139 1 1 3397993364 [913,1255,1198,1220] MNHN Chile 5 139 2 2