Achilia elfridae Raffray, 1904: 139 Jeannel, 1962: 426 Achilia picea Raffray, 1904: 139 Jeannel, 1962: 426-427 A revision of the Chilean Brachyglutini - Part 7. Revision of Achilia Reitter, 1890: A. cosmoptera species group (Coleoptera: Staphylinidae: Pselaphinae) Kurbatov, Sergey A. Cuccodoro, Giulio Sabella, Giorgio Revue suisse de Zoologie 2021 2021-04-23 128 1 135 156 977N Raffray, 1904 Raffray 1904 [968,1299,500,524] Insecta Staphylinidae Achilia Animalia Coleoptera 9 143 Arthropoda species elfridae   Figs 4, 23, 28, 49-51, 53, 55, 64      Achilia elfridae Raffray, 1904: 139, fig. 54 (head and antenna of male). –  Jeannel, 1962: 426, figs 190 (head), 191 (aedeagus).     Achilia picea Raffray, 1904: 139. –  Jeannel, 1962: 426-427( syn. nov.).   Type material ( 7 ex.): SOUTHERN CHILE: Región Los Ríos: Valdivia Prov.: MNHN; 1 ♂( lectotypeof  A. elfridae, here designated); labels verbatim “Lectotype / Type /  Bryaxis elfridaem.; Valdivia / Museum Paris; 1917; coll. Raffray/  A. elfridae; A. Raffraydet.”. – MNHN; 3 ♂and 2 ♀( Paralectotypesof  A. elfridae, here designated); labels verbatim “Paralectotype / Chili”. – MNHN; 1 ♀( holotypeof  Achilia picea); labels verbatim “Type/ Valdivia, var. valdiviensis; / Museum Paris; 1917; coll. Raffray/  A. picea; A. Raffraydet. /  piceaRaff.(handwritten by Jeannel)”.  Figs 7-15. Aedeagi (7, 8, 13), their internal sacs (9-12), and profemora (14, 15) of  Achilia. (7, 14)  A. quarantena. (8-12, 15)  A. angularis, specimens from Malleco, Nahuelbuta National Park. (10)  A. angularis, typefrom Chillan. (13)  A. pandemica.  Additional material examined ( 67 ex.): SOUTHERN ARGENTINA: Rio Negro Prov.: MHNG; 1 ♂; El Bolsón, Topal, n. 51;  06.II.1961. – SOUTHERN AND CENTRAL CHILE: Región Los Lagos: Llanquihue Prov.: MHNG; 28 ♂; Frutillar Bajo, Universidad Chile Forest Reserve;  100 m;  22.XII.1984/  02.II.1985; FIT, ravine mixed forest; S. & J. Peck. – Chiloé Prov.: MSNG; 1 ♂; Piruquina;  22.II.1991; site TC- 280; T. Cekalovic. – Región Los Ríos: Valdivia Prov.: PCTS; 1 ♂; Corral, 39° 95”S 73° 20”W;  06.XII.2013; car net. – PCVB; 1 ♂and 1 ♀; Panguipulli. – FMNH(FMHD #97-18); 2 ♂; Rincón de La Piedra, turnoff,  14.8 kmSE Valdivia; 39° 55’ 32’’S 73° 06’ 27’’W;  50 m;  11.I-01.II.1997; disturbed Valdivian rainforest, with  Nothofagus dombeyiand  Podocarpus saligna, flight intercept (windows) trap; A. Newton & M. Thayer990. – Región Araucanía: Cautín Prov.: UNHC; 1 ♂; Volcàn Villarica;  1120 m; site 654;  15-29.XII.1982;   Nothofagus dombeyiand  Saxegotheaforest with  Drimys, flight intercept (windows) trap; A. Newton & M. Thayer. – Malleco Prov.: MHNG; 1 ♂; Purén, Contulmo Natural Monument; station 17;  350 m;  11.XII.1984; S. & J. Peck. – MHNG; 2 ♂; same locality;  11.XII.1984-  13. II.1985; S. & J. Peck85-16. – MHNG; 3 ♂; same locality;  13.II.1985; mixed forest litter; S. & J. Peck. – MHNG; 1 ♂;  40 kmW CuraCautín;  1500 m,  12.XII.1984/  16.II.1985;  Notophagusand  Araucariaforest, malaise trap; S. & J. Peck. – FMNH(FMHD #85-905, #85-19); 1 ♂; Curaracautín(  40 kmE);  1500 m;  12.XII.1984/  16.II.1985;  Nothofagus-Araucariaforest, malaise trap; S. & J. Peck#85-19. – MHNG; 1 ♂; E Malalcahuello, 38° 26,0’S 71° 30,2’W;  1200 m;  11.II.2018; litter near stream; S. Kurbatov. – FMNH(FMHD #2002-041); 2 ♂; Nahuelbuta National Park, E of Guarderia Pichinahuel; 37° 48.20’S 73° 01.41’W;  1290 m;  05-24.XII.2002;  Araucaria- Nothofagus dombeyiwith  Chusqueabamboo, flight intercept trap; A. Newton, M. Thayer, A. Solodovnikov; D. J. Clarke & M. Chani1054. – Región Bío Bío: Ñuble Prov.: MNHN; 3 ♂and 7 ♀; Chillàn. – MNHS; 4 ♂and 5 ♀; Chillàn; P. Germain. – Región Maule: Curicó Prov.: MHNG; 1 ♂;  30 kmSE Curicó; 35° 13.3’S 70° 59.3’W;  200 m;  08.II.2018; litter near stream; S. Kurbatov.   Description:Body 1.25-1.35 mmlong, brown with reddish elytra, generally with darker head and sometimes also darker pronotum, some specimens brown with darker abdomen; antennae and legs reddish; palpi reddish or yellowish.  Male: Head as in Figs 51, 53, 55, wide, with occipital region slightly raised; frons flattened with enlarged and carinate sides on anterior half, and shallow triangular impression at middle; frons separated from enlarged frontal lobe by wide transverse sulcus. Antennae with scape and pedicel longer than wide; antennomere III longer than wide; remaining funicular articles slightly varying from antennomeres IV-VII slightly longer than wide to distinctly longer than wide and thinner, and from antennomere VIII wider than long to as long as wide; antennomere IX slightly wider than VIII and wider than long; antennomere X wider than long, and slightly wider than IX; antennomere XI elongate, about as long as VII-X combined. Metaventrite with deep and wide transverse sulcus on apical two-thirds, sulcus with prominent margins. First abdominal sternite slightly raised and flattened at middle ( Fig. 50); remaining abdominal sternites distinctly flattened at middle. Protibiae ( Fig. 23) with distal half swollen, distal third densely pubescent; mesotibiae ( Fig. 28) with distal half swollen and densely pubescent. Aedeagus ( Fig. 4) 0.24-0.26 mmlong, with suboval dorsal plate narrowed and frayed at apex; dorsal longitudinal struts divergent. Parameres relatively wide with long recurved seta on well-developed outer lobe; apical portion of parameres prolonged laterally as long tip; apex bearing ventral medial seta. Copulatory pieces consisting of pair of subequal sclerites curved at both ends and more robust basally.  Female: Similar to male except head not modified; antennae shorter with antennomere XI shorter; metaventrite, abdominal sternites, and legs unmodified.   Collecting data:Collected from November to February in different typeof forests (mixed forest, disturbed Valdivian rainforest,  Nothofagus dombeyiand  Saxegotheaforest with  Drimys, Nothophagusand  Araucariaforest, Araucaria-Nothofagus dombeyiwith  Chusqueabamboo), at elevations ranging from 50 mto 1290 m. Some specimens come from sifted samples of leaf and log litter, but most males have been collected by flight intercept traps, window traps, car netting, and malaise traps.   Distribution:  Achilia elfridaeis distributed in Southern Argentina( Rio Negro Province) and Southern and Central Chile(from Llanquihue to Curicó Provinces) ( Fig. 64: red circles).   Comments: Raffray (1904: 139)described  A. elfridaebased on an unspecified number of specimens from Chile, without further indication of locality, claiming to use the unpublished name attributed by Reitter to the specimens in his collection (now in Raffray’s collection). Jeannel (1962: 426-427)affirmed that the type of  Achilia elfridae Raffray, 1904collected in Valdivia was housed in the MNHN collections. In the MNHN we found in the Raffray collection a series of 6 specimensidentified by Raffray as  Achilia elfridae; the first specimen is a male labeled “Type /  Bryaxis elfridaem.; Valdivia / Museum Paris; 1917; coll. Raffray /  A. elfridae; A. Raffray det.” that we designate here as the lectotypeof  Achilia elfridae Raffray, 1904, and the remaining 5 specimens( 4 malesand 1 female) labeled “ Chili” without further label data are designated as paralectotypes.  Figs 23-32. Male protibiae (23-25), mesotibiae (26-31) and metatibiae (32) of  Achilia. (23, 28)  A. elfridae. (24, 30, 32)  A. angularis. (25, 31)  A. quarantena. (26)  A. covidia. (27)  A. blanchardi. (29)  A. pandemica.  Raffray (1904: 136)described  Achilia piceaon the basis of one femalethat Reitter had in his collection (now in Raffray’s collection) and placed it as a variety of  Bryaxis valdiviensis Reitter, 1883(=  Achilia blanchardi Raffray, 1904). According to Raffray (1904: 136)  A. piceadiffers from  A. blanchardiby having a longer head, thinner antennae, longer elytrae with denser punctures, and a darker body color. According to Jeannel (1962: 427)  A. piceastrongly resembles the females of  A. elfridae, from which it differs only by the darker coloration, the more transverse pronotum, and the narrower lateral bulges of the frontal lobe. We examined the holotypeand only known specimen of  A. picea; it is a female which indeed has a darker body color, but is otherwise similar in all other aspects to the females of  A. elfridae, and therefore we here place  A. picea Raffray, 1904as a junior synonym of  A. elfridae Raffray, 1904( syn. nov.). The males of  A. elfridaeare easily distinguished from other species of the  A. cosmopteragroup by the shape of the head ( Figs 51, 53, 55), and aedeagus ( Fig. 4). For characters to distinguish the females of  A. elfridaefrom those of  A. cosmopteraand  A. blanchardisee the “Comments” section for the two latter species. 3397993407 MNHN Raffray Chile A. Raffray 10 144 Valdivia Prov. Valdivia Prov. 9 143 1 1 Region Los Rios lectotype 3397993341 MNHN Chile 10 144 5 2 3 paralectotype 3397993307 MNHN Raffray Chile Jeannel A. Raffray Type Museum Paris 10 144 1 1 holotype 3397993306 1961-02-06 MHNG Argentina El Bolson, Topal Rio Negro Prov. 10 144 1 1 3397993311 ravine mixed forest 1984-12-22 1985-02-02 1984-12-22 MHNG S. & J. Peck Chile 100 Frutillar Bajo Llanquihue Prov 10 144 28 28 Region Los Lagos 3397993442 1991-02-22 MSNG T. Cekalovic Piruquina Chiloe Prov 10 144 1 1 3397993302 2013-12-06 car net PCTS Valdivia Prov Valdivia Prov 10 144 1 1 Region Los Rios 3397993376 PCVB Panguipulli 10 144 2 1 1 3397993427 disturbed Valdivian rainforest, with Nothofagus dombeyi and Podocarpus saligna 1997-01-11 1997-02-01 1997-01-11 flight intercept (windows) trap FMNH A. Newton & M. Thayer 50 -39.925556 Rincon de La Piedra 19 -73.1075 Rincon de La Piedra 10 144 2 2 3397993461 Nothofagus dombeyi and Saxegothea forest with Drimys 1982-12-15 1982-12-29 1982-12-15 flight intercept (windows) trap UNHC A. Newton & M. Thayer 1120 Volcan Villarica Cautin Prov 10 144 1 1 Region Araucania 3397993465 1984-12-11 MHNG S. & J. Peck 350 Puren Malleco Prov 10 144 1 1 3397993334 1984-12-11 1985-02-13 1984-12-11 MHNG S. & J. Peck Purén, Contulmo Natural Monument; station Malleco Prov. 10 144 2 2 3397993395 mixed forest litter 1985-02-13 MHNG S. & J. Peck Purén, Contulmo Natural Monument; station Malleco Prov. 10 144 3 3 3397993425 Notophagus and Araucaria forest 1984-12-12 1985-02-16 1984-12-12 malaise trap MHNG S. & J. Peck 1500 10 144 1 1 3397993394 Nothofagus-Araucaria forest 1984-12-12 1985-02-16 1984-12-12 malaise trap FMNH S. & J. Peck 1500 Curaracautin Curaracautin 10 144 1 1 3397993337 litter near stream 2018-02-11 MHNG S. Kurbatov 1200 -38.433334 E Malalcahuello 117 -71.503334 10 144 1 1 3397993339 Araucaria- Nothofagus dombeyi with Chusquea bamboo 2002-12-05 2002-12-24 2002-12-05 flight intercept trap FMNH A. Newton & M. Thayer & A. Solodovnikov & D. J. Clarke & M. Chani 1290 -37.803333 E of Guarderia Pichinahuel 11 -73.0235 Nahuelbuta National Park 10 144 2 2 3397993396 MNHN Nuble Prov. Nuble Prov. 10 144 10 7 3 Region Bio Bio 3397993345 [1005,1437,871,893] MNHS P. Germain Chillan 10 144 9 5 4 3397993393 litter near stream 2018-02-08 MHNG S. Kurbatov 200 -35.22167 Curico 119 -70.988335 Curico Prov 11 145 1 1 Region Maule