Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species López-García, Margarita M. Deloya, Cuauhtémoc The Coleopterists Bulletin 2019 2019-03-25 73 1 127 141 [408,502,1022,1045] Insecta Dynastidae Tomarus Animalia Coleoptera 9 135 Arthropoda genus    1. Pronotum without fovea or tubercle .......... 2  1 ʹ. Pronotum with fovea and tubercle, occasionally very small. If fovea absent, then pronotum nearly smooth, with minute, sparse punctures .......................................... 9   2. Apex of metatibia crenulate ( Fig. 24). Male protarsal inner claw wide and bifid, protarsomere 5 enlarged ........................... 3  2 ʹ. Apex of metatibia entire ( Fig. 25). Male protarsal inner claw protarsus simple, protarsomere 5 subcylindrical .................... 5   3. Protibia with 5 teeth: 3 large and 2 small denticules (one basal and one before second large tooth) ( Fig. 26). Mexicoand USA.... ................   Tomarus sallaei(Bates, 1888)  3 ʹ. Protibia with 4 teeth: 3 large and a small basal denticule ( Fig. 27) .............................. 4   4. Pygidium with dense rugosity on entire surface. Apex of metatibia with 9–10 spinules. Parameres short, with an acute lateral tooth on each side (Fig. 45). USAand Canada........ ...................   Tomarus relictus(Say, 1825)  4 ʹ. Pygidium with large, sparse punctures. Apex of metatibia with 14–15 spinules. Parameres long, without acute lateral tooth (Fig. 46). Ecuadorand Peru................................. ......   Tomarus peruvianus(Endrödi, 1970)   5. First apical tooth of mandible acute ( Fig. 28). Frontoclypeal region with a thick, straight carina ( Fig. 30). Mesotibia strongly narrowed before apex. Mexicoand Central America ...................   Tomarus nasutus(Burmeister, 1847)  5 ʹ. First apical tooth of mandible truncate ( Fig. 29). Frontoclypeal region with a pair of cariniform tubercles ( Fig. 31). Mesotibia not or only slightly contracted before apex ................. 6   6. Parameres with a large, dorsal tooth on each side, apex straight (Fig. 47). Females identifiable only by association with males. El Salvador, West Indies, and South America... ...............   Tomarus fossor(Latreille, 1813)  6 ʹ. Parameres without dorsal teeth, with a ventral tooth on each side, apex variable ........... 7   7. Parameres with acute, triangular apex ( Figs. 18–19). Metatibia strongly narrowed before apex. Apex of metatibia with 19 spinules. Brazil...................   Tomarus paranaensisLópez-Garc´ıa and Deloya, new species  7 ʹ. Parameres with apex truncate to rounded (Figs. 46–47). Metatibia not narrowed before apex, sides nearly parallel. Apex of metatibia with 25–29 spinules ............................ 8   8. Parameres wide, with parallel sides and minute spines on apicolateral third; ventral teeth short, triangular (Fig. 48). Females identifiable only by association with males. Argentina........... .................   Tomarus spinipenisNeita and Ratcliffe, 2017 Scutellum with sparse punctures. Apex of metatibia with 9–11 spinules. Bahamasand Cayman Islands.................................... ..................   Tomarus adoceteusRatcliffe and Cave, 2010 Brazil Rio de Janeiro 12 138 1 Rio de Janeiro