González-Sponga, 2005 : 106 Stenosfemuraia Bruvo-Mađarić et al . 2005 : 663 Astrin et al . 2006 : 445 Revision of the Venezuelan spider genus Stenosfemuraia González-Sponga, with new generic and specific synonymies (Araneae, Pholcidae) Huber, Bernhard A. Arias, Quintin Zootaxa 2017 2017-11-02 4341 4 495 508 524ZX Gonzalez-Sponga, 2005 Gonzalez-Sponga 2005 [151,753,1196,1222] Arachnida Pholcidae Stenosfemuraia GBIF Animalia Araneae 5 500 Arthropoda species cuadrata     Stenosfemuraia cuadrata  González-Sponga, 2005: 106, pl. 4, figs 1–9 ( ♂ ♀).  Stenosfemuraiasp.:  Bruvo-Mađarić et al. 2005: 663(DNA sequence: 28S);  Astrin et al. 2006: 445(DNA sequences: 16S, CO1).   Diagnosis.Males are easily distinguished from  S. parvaby armature of male chelicerae ( Fig. 24; one pair of simple frontal apophyses; see also fig. 3 in González-Sponga 2005); from  S. pilosaby shape of procursus ( Figs 47–48; slender in dorsal view, without subdistal side branch, with simple tip; possibly indistinguishable from  S. parva); from both species also by more slender femora (~0.13 versus 0.19–0.21 in  S. parvaand 0.29–0.34 in  S. pilosa). Females differ from congeners by sclerotized lateral elements of epigynal plate ( Figs 19, 26; absent in  S. pilosa; rather parallel orientation in  S. parva) and by distinctive shape of anteriorly contiguous pore plates ( Figs 21, 27). Males and females also differ from congeners by longer legs ( ♂tibia 1 L/d: ~60 versus<40; ♂tibia 1:>5.0 versus<4.0; ♀tibia 1:>3.0 versus<3.0).     Typematerial. VENEZUELA:  Vargas: 22♂ 16♀ types(+8 juvs and 4 prosomata) (see Notesbelow), MIZA( GS 1164part), Hacienda El Limón, Municipio Carayaca,  950 m[~ 10.532°N, 67.119°W; see Notes below],  12.viii.1989–   4.v.1991( A.R. Delgado de González, M.A. González-Sponga), 3♂examined by BAH.    Othermaterial examined. VENEZUELA:  Aragua: 4♂ 3♀ 1 juv., ZFMK(Ar 18253), forest above Colonia Tovar ( 10.417°N, 67.300°W), ~  2100 ma.s.l., under dead leaves on ground,  26.xi.2002( B.A. Huber);  4♂ 6♀in pure ethanol, ZFMK( Ven02/100-10), same data. 2♀, MNHN (Ar 10052 part), with Simon’s label “14653 Psil.— Tovar!”, no further data [leg. E. Simon, 1887–88].   Redescription. Male(ZFMK Ar 18253). MEASUREMENTS. Total body length 2.5, carapace width 1.0. Distance PME-PME 80 µm, diameter PME 100 µm, distance PME-ALE 90 µm, distance AME-AME 20 µm, diameter AME 20 µm. Sternum width/length: 0.75/0.50. Leg 1: 21.1 (4.9 + 0.4 + 5.1 + 8.8 + 1.9), tibia 2: 3.1, tibia 3: 2.4, tibia 4: 2.8; tibia 1 L/d: 60. Femora 1–4 width (at half length): all ~0.13. COLOR (in ethanol). Carapace ochre-yellow with brown marginal bands and median mark including ocular area and clypeus ( Fig. 15); sternum whitish; legs ochre-yellow, femora and tibiae with subdistal darker rings (and indistinct light tips); abdomen pale gray, dorsally and laterally with dark bluish internal marks, ventrally with indistinct light brown plate in front of gonopore, without mark behind gonopore. BODY. Habitus as in Figs 15–16; ocular area raised; carapace elevated, with deep median furrow; clypeus and sternum unmodified. CHELICERAE. With one pair of simple (undivided) frontal apophyses ( Fig. 24). PALPS. As in Figs 22–23; coxa with retrolatero-ventral apophysis; trochanter barely modified; femur with retrolatero-ventral process proximally and small ventral apophysis distally; procursus very simple, slender in dorsal view, without subdistal side branch ( Figs 47–48); bulb with two dorsal processes, distal apophysis curved, very slender towards tip ( Figs 45–46). LEGS. Without spines; with curved hairs on metatarsi 1–3, few vertical hairs; retrolateral trichobothrium on tibia 1 at 6%; prolateral trichobothrium present on tibia 1; tarsus 1 with ~25 pseudosegments, distally distinct.  Male(variation). Tibia 1 in8 other males: 5.0–5.7 (mean 5.3). Internal abdominal marks less distinct in types. One male with few curved hairs also on tibiae 1–3.   FIGURES 15–21.  Stenosfemuraia cuadrata, ZFMK Ar 18253. 15–18. Male and female, dorsal and lateral views. 19–20. Epigynum, ventral and lateral views. 21. Cleared female genitalia, dorsal view (arrows: lateral sclerites/‘wings’).   FIGURES 22–27.  Stenosfemuraia cuadrata, ZFMK Ar 18253. 22–23. Left pedipalp, prolateral and retrolateral views. 24. Male chelicerae, frontal view. 25–26. Epigynum, lateral and ventral views. 27. Cleared female genitalia, dorsal view. Scale lines: 0.3.  Female.In general similar to male ( Figs 17–18), legs without curved hairs. Tibia 1 in 7 females: 3.3–3.7 (mean 3.5). Epigynum as in Figs 19–20, 25–26; with large protruding weakly sclerotized area in front of light brown epigynal plate, with pair of dark lateral sclerites in transversal orientation, posterior margin evenly curved; posterior plate simple. Internal genitalia as in Figs 21and 27, with pair of anteriorly contiguous pore-plates, membranous median sac, and anteriorly diverging sclerites/‘wings’.  Natural history.All specimens were collected from forest leaf litter. González-Sponga (2005)specifically mentions  Cecropialeaves.   Distribution.Known from medium to high elevation forests ( 950–2100 ma.s.l.) in Aragua(Tovar area) and Vargasstates, Venezuela( Fig. 1).  Notes. González-Sponga (2005)reported 1♂ holotype(“1164a”), 1♀ paratype(“1164b”), and 21♂ 27♀adult paratypeswithout specific collection number. Only one vial was found at MIZA, labeled “1164” and containing 22♂ 16♀8 juvs and 4 prosomata. Of the 22♂, three belong to  S. parvaand were separated (see above) (and clearly labeled as being part of the  S. cuadrata typeseries). The label says “Hda El Limón, Dpto Vargas, DF” (which differs only with respect to administrative units from the locality as published in the original description: “Hacienda El Limón, Municipio Carayaca, Estado Vargas) and lists four dates: “ 12-8-89, 29-10-89, 10-8-90, 4-5- 91” (which agrees with the range in the original description). We suspect that this is the entire typeseries and that a holotypewas never physically separated from this series. Since the identity of the species is beyond doubt, and since the holotypemight just be misplaced rather than lost, we prefer not to designate a neotypeand to treat the typespecimens unspecifically as “ types”.  González-Sponga (2005)gives the distribution of  S. cuadrataas “Galipán, Parque Nacional El Avila, Estado Vargas”. This does not agree with the typelocality. The exact coordinates of the typelocality are not known to us, but Hacienda El Limón is close to the town of Carayaca, while Galipán [San Antonio de Galipán] is a town within the limits of El Ávila National Park and lies about 25 kmE of the typelocality. We do not have an explanation for this contradiction. In addition, the coordinates in González-Sponga (2005)are clearly wrong (about 32 kmSW of Carayaca, in the state of Aragua). Some measurements in González-Sponga (2005)are clearly wrong, e.g.: female carapace width (0.4; should be ~0.7). 1677458077 1989-08-12 MIZA Hacienda El Limon & Municipio Carayaca Venezuela 950 10.532 Notes 77 -67.119 5 500 GS 1164 38 16 22 Vargas holotype 1677458072 [302,1272,1790,1815] 1991-05-04 BAH A. R. Delgado de Gonzalez & M. A. Gonzalez-Sponga Venezuela Delgado de Gonzalez 5 500 3 3 Vargas holotype 1677458071 2002-11-26 ZFMK B. A. Huber Venezuela 2100 10.417 Other 77 -67.3 5 500 8 3 4 Aragua 1677458073 2002-11-26 ZFMK Ven Venezuela 2100 10.417 Other 77 -67.3 5 500 10 6 4 Aragua