A new species, new genus and new records of heterostigmatic mites (Acari: Heterostigmata) phoretic on scarab beetles of the subfamily Orphninae (Coleoptera: Scarabaeidae) Khaustov, Alexander A. Frolov, Andrey V. Zootaxa 2018 2018-11-07 4514 2 181 201 Paoli, 1911 Paoli 1911 [238,540,221,247] Arachnida Scutacaridae Pygmodispus GBIF Animalia Prostigmata 18 199 Arthropoda genus   Typespecies:  Pygmodispus equestris Paoli, 1911, by original designation.