Herbst, 1791: 25 Stimpson 1858: 72 Rankin 1900: 533 Schmitt 1935: 198 Provenzano 1959: 363 Forest & de Saint Laurent 1968: 106 Narchi & Hebling 1972: 67 Sánchez & Campos 1978: 22 Coelho & Ramos-Porto 1986: 52 Campos & Lemaitre 1994: 147 Rieger 1998: 422 Melo 1999: 42 Souza & Serejo 2007: 341 Almeida et al . 2010: 335 McLaughlin et al . 2010: 19 Lemaitre & Tavares 2015: 453 Tavares et al. 2017: 171 Poupin 2018: 154 Pagurus tibicen Bosc 1802: 78 Milne Edwards, 1836: 279 1848: 64 Calcinus sulcatus Stimpson 1858: 72 Smith 1869: 17 Rathbun 1900: 144 Moreira 1901: 27 Neumann, 1878: 31 Alcock 1905: 164 Gordan 1956: 304 Morgan 1991: 907 Paguroids (Decapoda: Anomura: Diogenidae and Paguridae) of the remote oceanic Archipelago Trindade and Martin Vaz, off southeast Brazil, with new records, description of three new species and zoogeographical notes Lima, Daniel Tavares, Marcos Jr, Joel Braga De Mendonça Zootaxa 2019 2019-11-05 4694 1 1 63 189f2395-b631-42dd-8ce8-39a952c990e9 (Herbst, 1791) Herbst 1791 [151,534,1047,1074] Malacostraca Diogenidae Calcinus Animalia Decapoda 6 7 Arthropoda species tibicen  ( Fig. 3A–D)     Cancer tibicen  Herbst, 1791: 25, pl. 23, fig. 7;  Stimpson 1858: 72;  Rankin 1900: 533, pl. 17, fig. 1;  Schmitt 1935: 198, fig. 60a, b;  Provenzano 1959: 363, fig. 4; 1960: 120; 1961: 152;  Forest & de Saint Laurent 1968: 106;  Narchi & Hebling 1972: 67, fig. 1;  Sánchez & Campos 1978: 22, fig. 5;  Coelho & Ramos-Porto 1986: 52;  Campos & Lemaitre 1994: 147, figs. 1c–4c, 5d, 6c, 7c, f, I;  Rieger 1998: 422;  Melo 1999: 42, fig. 2;  Souza & Serejo 2007: 341, fig. 2;  Almeida et al. 2010: 335;  McLaughlin et al. 2010: 19;  Lemaitre & Tavares 2015: 453, tab. 1;  Tavares et al.2017: 171, tab. 1;  Poupin 2018: 154.    Pagurus tibicen.—  Bosc 1802: 78.    Pagurus sulcatusH.  Milne Edwards, 1836: 279;  1848: 64.    Calcinus sulcatus.—  Stimpson 1858: 72;   Smith1869: 17, 39;  Rathbun 1900: 144; Verril 1900: 578; Benedict 1901a: 141, pl. 5, figs. 3, 3a;  Moreira 1901: 27, 86; 1920: 132; Verril 1908: 439, figs. 56, 57, pl. 28, fig. 7.    Calcinus formosus  Neumann, 1878: 31;  Alcock 1905: 164;  Gordan 1956: 304;  Morgan 1991: 907.   Trindade and Martin Vaz specimens.  2 males( MZUSP 36068), Brazil, Espírito Santo, Martin Vaz, 20°28’26.98”S, 28°51’20.98”W, C.H. Guimarãescoll.,  22.i.2012, SCUBA,  19.2 m;  2 males( MZUSP 36135), idem, Praia dos Andradas, 20°30’45.7”S, 29°18’21.9”W, J.B. Mendonçacoll.,  24.vii.2013, SCUBA, 12.3 mm;  1 female( MZUSP 36163), Brazil, Espírito Santo, Trindade Island, Ponta Norte, 20°29’18.7”S, 29°20’18.3”W, J.B. Mendonçacoll.,  23.vi.2016, SCUBA,  11.3 m;  2 males, 1 female( MZUSP 36084), idem, Ponta Norte, 20°29’18.7”S, 29°20’18.3”W, J.B. Mendonçacoll.,  23.vi.2016, SCUBA,  11.3 m;  4 males, 3 females, 2 ovigerous females ( MZUSP 36062), idem, Enseada Crista do Galo, 20°29’08.7”S, 29°20’10.7”W, C.H. Guimarãescoll.  07.ii.2012, SCUBA,  22.2 m;  3 males, 1 female( MZUSP 36170), idem, Crista do Galo, 20°29’22.1”S, 29°20’03.1”W, J.B. Mendonçacoll.,  17.vii.2012, SCUBA,  2.5 m;  1 male, 1 female( MZUSP 36088), idem, Praia dos Cabritos, 20°29’32.0”S, 29°19’46.5”W, J.B. Mendonçacoll.,  14.vi.2012, SCUBA,  9.4 m;  2 males, 1 female( MZUSP 36127), idem, Enseada das Orelhas, 20°29’32.3”S, 29°20’32.6”W, J.B. Mendonçacoll.,  25.vi.2012, SCUBA,  14.9 m;  1 female( MZUSP 36169), idem, Enseada de Orelhas, 20°29’40.2”S, 29°20’32.9”W, J.B. Mendonçacoll.,  06.vii.2013, SCUBA,  14 m;  3 males, 1 female, 6 ovigerous females ( MZUSP 36118), idem, Enseada das Orelhas, 20°29’40.2”S, 29°20’32.9”W, J.B. Mendonçacoll.,  01.viii.2015, SCUBA,  8.9 m;  9 male, 1 ovigerous female( MZUSP 36120), idem, Enseada das Orelhas, 20°29’40.2”S, 20°20’32.9”W, J.B. Mendonçacoll.,  01.viii.2015, SCUBA,  8.9 m;  3 males, 2 females( MZUSP 36063) idem, Ponta do Monumento, 20°30’10.3”S, 29°20’36.1”W, J.B. Mendonçacoll.,  02.iv.2014, SCUBA,  8.4 m;  1 male( MZUSP 36087), idem, Enseada das Orelhas, Simone et al., coll.,  24.vii.2011;  7 males, 3 females, 1 ovigerous female( MZUSP 36108), idem, Enseada das Orelhas, J.B. Mendonçacoll.,  01.viii.2015, SCUBA,  8.9 m;  1 male, 1 ovigerous female( MZUSP 36119), idem, Laje Enseada Noroeste, 20°29’51.0”S, 29°20’44.3”W, C.H. Guimarãescoll.,  14.ii.2012, SCUBA,  20 m;  1 male, 2 females( MZUSP 36104), Laje da Ponta Noroeste, 20°29’57.8”S, 29°20’39.2”W, J.B. Mendonçacoll.,  09.vii.2012, SCUBA,  17.7 m;  1 male( MZUSP 36064), idem, Enseada dos Portugueses, stn Farol, 20°29’52.3”S, 29°19’15.6”W, J.B. Mendonçacoll.,  20.v.2014, SCUBA,  13.1 m;  1 female( MZUSP 36171), idem, Enseada dos Portugueses, stn Farol, J.B. Mendonçacoll.,  08.vii.2015, artificial reef substrate;  2 females( MZUSP 36076), idem, Enseada dos Portugueses, stn Farol, 20°29’52.3”S, 29°19’15.6”W, J.B. Mendonçacoll.,  24.vi.2016, SCUBA,  11.3 m;  1 male, 2 females( MZUSP 36130), idem, Enseada dos Portugueses, stn Farol, 20°29’52.3”S, 29°19’15.6”W, J.B. Mendonçacoll.,  10.vii.2012, SCUBA,  13.2 m;  2 females( MZUSP 36123), idem, Enseada dos Portugueses, stn SECON, 20°30’20.9”S, 29°18’43.7”W, J.B. Mendonçacoll.,  18.vii.2013, SCUBA,  12.2 m;  2 males, 2 females, 2 ovigerous females ( MZUSP 36103), Praia dos Portugueses, stn Rampa Nova, 20°30’17.7”S, 29°18’56.7”W, J.B. Mendonçacoll.,  23.iv.2014, SCUBA,  10.7 m;  1 male, 1 ovigerous female( MZUSP 36092), idem, Ponta do Monumento, 20°30’10.3”S, 29°20’36.1”W, J.B. Mendonçacoll.,  13.vii.2012, SCUBA,  8.1 m;  1 male( MZUSP 36082), idem, Ponta da Calheta, Abbate et al., coll.,  23.vii.2011, SCUBA;  1 male( MZUSP 36078), idem, Ponta da Calheta, J.B. Mendonçacoll.,  03.viii.2015, SCUBA,  17.7 m;  2 males( MZUSP 36101), idem, Ponta da Calheta, 20°30’18.7”S, 29°18’31.6”W, J.B. Mendonçacoll.,  20.v.2014, SCUBA,  16.3 m;  1 male( MZUSP 36098), idem, Praia da Calheta, stn ECIT, 20°30’19.5”S, 29°18’47.3”W, J.B. Mendonçacoll.,  26.vi.2012, SCUBA,  14.3 m;  1 ovigerous female( MZUSP 36093), idem, Praia da Calheta, stn SECON, 20°30’26.1”S, 29°18’44.2”W, J.B. Mendonçacoll.,  03.vii.2012, SCUBA;  2 males, 1 female( MZUSP 36072), idem, Praia da Calheta, 20°30’29.5”S, 29°18’37.0”W, J.B. Mendonçacoll.,  21.vi.2016, SCUBA,  12.5 m;  1 female, 1 ovigerous female( MZUSP 36106), idem, Praia da Calheta, 23°49’10.7”S, 29°24’17.6”W, V.S. Amaral& D. Abbatecoll.,  27.iii.2011;  3 males, 1 female, 3 ovigerous females ( MZUSP 36126), idem, Praia da Calheta, 20°30’29.5”S, 29°18’37.0”W, J.B. Mendonçacoll.,  10.iv.2014, snorkeling,  5 m;  2 males( MZUSP 36125), idem, Ilha da Racha, 20°30’26.5”S, 29°20’48.0”W, J.B. Mendonçacoll.,  13.vii.2012, SCUBA,  23.8 m;  6 males, 1 female( MZUSP 36107), idem, Enseada da Cachoeira, Praia do Eme, 20°30’55.6”S, 29°20’21.7”W, J.B. Mendonçacoll.,  12.vii.2012, SCUBA;  3 males, 1 female( MZUSP 36131), idem, Enseada da Cachoeira, 20°30’57.1”S, 29°20’15.2”W, J.B. Mendonçacoll.,  09.vii.2012, SCUBA,  13.8 m;  1 male, 1 female, 1 ovigerous female( MZUSP 36100), idem, Praia dos Andradas, 20°28’47.69”S, 29°18’24.03”W, L.R. Simonecoll.,  06.ii.2012;  1 male, 2 females( MZUSP 36089), idem, Praia dos Andradas, 20°30’71.8”S, 29°18’24.7”W, J.B. Mendonçacoll.,  17.vii.2013, SCUBA,  9.9 m;  1 male, 2 females( MZUSP 36166), idem, Praia das Tartarugas, 20°30’54.1”S, 29°18’18.2”W, J.B. Mendonçacoll.,  05.vii.2013;  2 males( MZUSP 36096), idem, stn Parcel, 20°31’01.3”S, 29°17’56.9”W, J.B. Mendonçacoll.,  26.vi.2012, SCUBA,  9.5 m;  2 males( MZUSP 36111), idem, Parcel das Tartarugas, stn tide pool, 20°31’10.4”S, 29°17’58.4”W, J.B. Mendonçacoll.,  19.vi.2016, intertidal and snorkeling, 1.0 m;  1 juvenile( MZUSP 39876), idem, stn tide pool, 20°31’10.4”S, 29°17’58.4”W, J.B. Mendonçacoll.,  02.ii.2019, snorkeling,  0.5 m;  1 juvenile( MZUSP 39877), idem, stn tide pool, 20°31’10.4”S, 29°17’58.4”W, J.B. Mendonçacoll.,  15.xi.2017, 1.0 m;  2 males, 4 females, 11 juveniles( MZUSP 36164), idem, Praia do Parcel, 20°31’11.4”S, 29°18’00.6”W, J.B. Mendonçacoll.,  15.vii.2012, snorkeling;  3 females, 1 male( MZUSP 36091), idem, Praiado Príncipe, 20°31’20.1”S, 29°18’34.7”W, J.B. Mendonçacoll.,  19.vi.2012, SCUBA,  12.2 m;  3 males, 1 female( MZUSP 36117), idem, Enseadado Príncipe, stn Pedra da Garoupa, 20°31’35.5”S, 29°18’34.0”W, J.B. Mendonçacoll.,  16.vii.2013, SCUBA,  10.4 m;  8 males, 5 females( MZUSP 36133), idem, Enseada da Cachoeira, Farrilhões, 20°31’22.4”S, 29°19’52.0”W, J.B. Mendonçacoll.,  09.vii.2013, SCUBA,  10.4 m;  16 males, 7 females( MZUSP 36109), idem, Enseada da Cachoeira, Farrilhões, 20°31’22.4”S, 29°19’52.0”W, J.B. Mendonçacoll.,  20.vi.2012, SCUBA,  11.8 m;  1 male, 1 female, 1 ovigerous female( MZUSP 36136), idem, Enseada da Cachoeira, stn Farrilhões, 20°31’22.4”S, 29°19’52.0”W, J.B. Mendonçacoll.,  25.x.2014, SCUBA,  10.7 m;  1 female, 1 male( MZUSP 36162), idem, Praia do Lixo, stn Paredão, 20°31’29.8”S, 29°19’43.9”W, J.B. Mendonçacoll.,  22.iv.2014, SCUBA,  14.5 m;  1 male( MZUSP 36116), idem, Enseada do Lixo, 20°31’29.8”S, 29°19’43.9”W, J.B. Mendonçacoll.,  02.vii.2012, SCUBA,  25 m;  1 male( MZUSP 36075), idem, Enseada do Lixo, 20°31’43.5”S, 29°19’28.1”W, C.H. Guimarãescoll.,  17.ii.2012, SCUBA,  23 m.  Comparative material examined.  Calcinus tibicen:  5 males, 4 females, 1 juvenile( MZUSP 28061), Barbados, Christ Church, Turtle Beach, 13°04’30.77”N, 59°36’16.88”W, L.R.L Simone et al. coll.,  13.x.2012, snorkeling, 0–1.0 m;  1 male(28067), Grenada, Grand Anse Bay, 12°01’45.68”N, 61°46’18.67”W, L.R.L. Simone et al.   FIGURE 2. (A–F) Trindade Island. (A) mixed sand and gravel beach and tide pool. (B–F) sublittoral habitats at water depths of less than 30 m: (B, C) calcareous reefs with algal mats. (D) rocky and calcareous boulders of several sizes. (E) sand bottom with patches of macroalgae and sponges. (F) rhodolith beds. Photographs by A. Pol. (A) and J.B. Mendonça Jr. (B–F). coll., 08.x.2012, SCUBA,   0–10 m; 6 males, 1 ovigerous female(29753), Costa Rica, Puerto Viejo, 9°39’28”N, 82°45’20”W, G. Guzmáncoll.,  13.vii.2013, intertidal;  1 male( MZUSP 13894), Brazil, Pernambuco, Fernando de Noronha, Praia do Meio, Simone& Souzacoll.,  22.vii.1999;  1 male, 7 ovigerous females ( MZUSP 17712), Brazil, Bahia, Abrolhos, W. Narchicoll.,  ix.1965;  4 males, 3 ovigerous females ( MZUSP 7028), Brazil, Bahia, Abrolhos, Ilha Redonda, P. Youngcoll.,  21.i.1985;  1 male( MZUSP 34990), Brazil, Ceará, Fortaleza, Meirelesrock beach, Anker& Feitosacoll.,  15.vi.2011;  1 ovigerous female( MZUSP 29139), Brazil, Espírito Santo, Piúma, 20°50’57.8”S, 40°28’25.3”W, A. Jóriocoll.,  28.iii.2010,  21.5 m ; 2 males( MZUSP 33644), Brazil, Espírito Santo, Guarapari, Praia do Ermitão, R.C. Marquescoll.,  18.vii.2008;  1 ovigerous female( MZUSP 36746), Brazil, Rio de Janeiro, Baía de Guanabara, Forte de Santa Cruz, BIOPLAT-BG coll.,  05.i.2001;  2 males( MZUSP 8666), Brazil, Rio de Janeiro, Rio de Janeiro, Praia de Itaipú, W. Vergaracoll.,  10.i.1987, 2 m;  3 males( MZUSP 12991), Brazil, São Paulo, São Sebastião, Saco Grande ; 1 ovigerous female( MZUSP 25036), Brazil, São Paulo, Ilha Bela, Praia do Portinho, 23°49’S, 45°23’W, E.P. Gonçalves et al. coll.,  26–27.ii.2015,  0–3 m ; 5 males( MZUSP 9431), Brazil, Santa Catarina, Praia da Armação, W.L. Vergaracoll.,  16.vi. 1986, 1 m;  2 males( MZUSP 35066), Brazil, Santa Catarina,  03.xi.2011.   Typelocality.Unknown (see Schmitt 1935).   Distribution. Bermuda, Florida, Gulf of Mexico, Antilles, Panama, Colombia, Venezuela, and Brazil(from Cearáto Santa Catarina. Oceanic islands: Fernando de Noronha ( Nucci & Melo 2015), Rocas Atoll ( Paiva et al. 2007) and Archipelago Trindade and Martin Vaz, present study).   Remarks.Forty-five species are currently attributed to  Calcinus, 35 of which are from the Indo-West Pacific, four from the eastern Pacific, three from the eastern Atlantic, and three from the western Atlantic ( Poupin & Bouchard 2006; Malay et al. 2018). The specimens from TMV are assigned to  Calcinus tibicenon the basis of the smooth upper margin of the right chela, whereas in the other two western Atlantic species,  C. verrilli( Rathbun, 1901)and  C. urabaensis Campos & Lemaitre, 1994, it is tuberculated or spinous. Additionally, in  C. tibicenthe posterior margin of the telson left lobe is provided with a single spine, whereas in  C. verrilliand  C. urabaensisit is armed with several spines.   FIGURE 3. (A–D)  Calcinus tibicen( Herbst, 1791)from Trindade Island. (A, B) male sl 6.0 mm (MZUSP 36122). (C) juvenile sl 2.0 mm (MZUSP 39876). (D) juvenile sl 3.0 mm (MZUSP 39877). Note the differences in color pattern between specimens. Photographs by A. Pol. Scale bars: A–C, 10 mm; D, 5.0 mm.   Calcinus tibicenand the eastern Atlantic  C. talismaniA. Milne-Edwards & Bouvier, 1892, superficially resemble each other in having the upper margin of the right chela smooth, and grooved P3 propodus, but can be easily disguished in that the dorsal margin of the P3 propodus is distinctly angulous in  C. tibicen, whereas in  C. talismaniit is rounded (Poupin 2003).  Mandai et al.(2018a)considered the differences of color pattern among northeastern and southeastern Brazilian specimens of  C. tibicena case of polymorphism and, without further explanation, suggested that it “might be related to the kind of nutrition as well different ecological and evolutionary predation characteristics.” Later on, however, Mandai et al. (2018b), based on two mitochondrial markers, found that northwestern Atlantic specimens differed from northeastern and southeastern Brazilian coast specimens, despite the lack of morphological evidences to support each genetic group. A number of sibling species in  Calcinuscan be split almost exclusively by color pattern (see for instance Poupin & Lemaitre 2003). Live coloration in  C. tibicencan indeed vary dramatically, but color variants are found within the same population ( Fig. 3) and in a limited geographical area and, hence, do not match the genetic groups of Mandai et al.(2018b). 2446840894 2012-01-22 MZUSP C. H. Guimaraes Brazil 19 -20.474161 Martin Vaz 1 -28.855827 6 7 MZUSP 36068 2 2 Espirito Santo 2446840985 2013-07-24 MZUSP J. B. Mendonca Brazil 19 -20.512695 Praia dos Andradas 1 -29.306082 6 7 MZUSP 36135 2 2 Espirito Santo 2446840516 2016-06-23 MZUSP J. B. Mendonca Brazil 11 -20.488527 Ponta Norte 1 -29.338417 Trindade Island 6 7 MZUSP 36163 1 1 Espirito Santo 2446841232 2016-06-23 MZUSP J. B. Mendonca Brazil 11 -20.488527 Ponta Norte 1 -29.338417 Trindade Island 6 7 MZUSP 36084 3 1 2 Espirito Santo 2446841401 2012-02-07 MZUSP C. H. Guimaraes Brazil 22 -20.48575 Enseada Crista do Galo 1 -29.336306 Trindade Island 6 7 MZUSP 36062 7 3 4 Espirito Santo 2446840932 2012-07-17 MZUSP J. B. Mendonca Brazil 3 -20.489473 Crista do Galo 1 -29.334194 Trindade Island 6 7 MZUSP 36170 4 1 3 Espirito Santo 2446840844 2012-06-14 MZUSP J. B. Mendonca Brazil 9 -20.492222 Praia dos Cabritos 1 -29.329584 Trindade Island 6 7 MZUSP 36088 2 1 1 Espirito Santo 2446841426 2012-06-25 MZUSP Enseada das Orelhas & J. B. Mendonca Brazil 15 -20.492306 Praia dos Cabritos 1 -29.34239 Trindade Island 6 7 MZUSP 36127 3 1 2 Espirito Santo 2446841087 2013-07-06 MZUSP J. B. Mendonca Brazil 14 -20.494501 Enseada de Orelhas 1 -29.342472 Trindade Island 6 7 MZUSP 36169 1 1 Espirito Santo 2446840432 2015-08-01 MZUSP Enseada das Orelhas & J. B. Mendonca Brazil 9 -20.494501 Enseada de Orelhas 1 -29.342472 Trindade Island 6 7 MZUSP 36118 4 1 3 Espirito Santo 2446841294 2015-08-01 MZUSP Enseada das Orelhas & J. B. Mendonca Brazil 9 -20.494501 Enseada de Orelhas 1 -20.342472 Trindade Island 6 7 MZUSP 36120 9 9 Espirito Santo 2446841371 2014-04-02 MZUSP J. B. Mendonca Brazil 8 7 8 -20.502861 Ponta do Monumento 1 -29.34336 Trindade Island 6 7 MZUSP 36063 5 2 3 Espirito Santo 2446841217 [190,1175,189,216] 2011-07-24 MZUSP Enseada das Orelhas & Simone Brazil 8 -20.502861 Ponta do Monumento 1 -29.34336 Trindade Island 7 8 MZUSP 36087 1 1 Espirito Santo 2446840793 2015-08-01 MZUSP Enseada das Orelhas, J. B. & Mendonca Brazil 9 -20.502861 Ponta do Monumento 1 -29.34336 Trindade Island 7 8 MZUSP 36108 10 3 7 Espirito Santo 2446840943 2012-02-14 MZUSP C. H. Guimaraes Brazil 20 -20.4975 Laje Enseada Noroeste 1 -29.34564 Trindade Island 7 8 MZUSP 36119 1 1 Espirito Santo 2446841462 2012-07-09 MZUSP J. B. Mendonca Brazil 18 -20.49939 Laje da Ponta Noroeste 1 -29.344223 7 8 MZUSP 36104 3 2 1 Espirito Santo 2446841357 2014-05-20 MZUSP Enseada dos Portugueses & J. B. Mendonca Brazil 13 -20.497862 Farol 1 -29.321001 7 8 MZUSP 36064 1 1 Espirito Santo 2446841497 2015-07-08 MZUSP Enseada dos Portugueses & J. B. Mendonca Brazil 13 -20.497862 Farol 1 -29.321001 7 8 MZUSP 36171 1 1 Espirito Santo 2446841494 2016-06-24 MZUSP Enseada dos Portugueses & J. B. Mendonca Brazil 11 -20.497862 Farol 1 -29.321001 7 8 MZUSP 36076 2 2 Espirito Santo 2446840578 2012-07-10 MZUSP Enseada dos Portugueses & J. B. Mendonca Brazil 13 -20.497862 Farol 1 -29.321001 7 8 MZUSP 36130 3 2 1 Espirito Santo 2446841477 2013-07-18 MZUSP Enseada dos Portugueses & J. B. Mendonca Brazil 12 -20.505806 Farol 1 -29.312138 7 8 MZUSP 36123 2 2 Espirito Santo 2446840503 2014-04-23 MZUSP J. B. Mendonca Brazil 11 -20.504917 Rampa Nova 1 -29.31575 Praia dos Portugueses 7 8 MZUSP 36103 4 2 2 Espirito Santo 2446841270 2012-07-13 MZUSP J. B. Mendonca Brazil 8 -20.502861 Ponta do Monumento 1 -29.34336 Praia dos Portugueses 7 8 MZUSP 36092 1 1 Espirito Santo 2446841441 2011-07-23 MZUSP Abbate Brazil 8 -20.502861 Ponta da Calheta 1 -29.34336 Praia dos Portugueses 7 8 MZUSP 36082 1 1 Espirito Santo 2446841352 [262,1432,728,756] 2015-08-03 MZUSP J. B. Mendonca Brazil 18 -20.502861 Ponta da Calheta 1 -29.34336 Praia dos Portugueses 7 8 MZUSP 36078 1 1 Espirito Santo 2446841080 2014-05-20 MZUSP J. B. Mendonca Brazil 16 -20.505194 Ponta da Calheta 1 -29.308777 Praia dos Portugueses 7 8 MZUSP 36101 2 2 Espirito Santo 2446841436 2012-06-26 MZUSP J. B. Mendonca Brazil 14 -20.505417 Praia da Calheta 1 -29.313139 Praia dos Portugueses 7 8 MZUSP 36098 1 1 Espirito Santo 2446841329 2012-07-03 MZUSP J. B. Mendonca Brazil 14 -20.50725 Praia da Calheta 1 -29.312277 Praia dos Portugueses 7 8 MZUSP 36093 1 Espirito Santo 2446841170 2016-06-21 MZUSP J. B. Mendonca Brazil 13 -20.508194 Praia da Calheta 1 -29.310278 Praia dos Portugueses 7 8 MZUSP 36072 3 1 2 Espirito Santo 2446841425 2011-03-27 MZUSP V. S. Amaral & D. Abbate Brazil 13 -23.81964 Praia da Calheta 1 -29.404888 Praia dos Portugueses 7 8 MZUSP 36106 1 1 Espirito Santo 2446841456 2014-04-10 MZUSP J. B. Mendonca Brazil 5 -20.508194 Praia da Calheta 1 -29.310278 Praia dos Portugueses 7 8 MZUSP 36126 4 1 3 Espirito Santo 2446841502 2012-07-13 MZUSP J. B. Mendonca Brazil 24 -20.50736 Ilha da Racha 1 -29.346668 Praia dos Portugueses 7 8 MZUSP 36125 2 2 Espirito Santo 2446840923 2012-07-12 MZUSP J. B. Mendonca Brazil 24 -20.515444 Praia do Eme 1 -29.339361 Enseada da Cachoeira 7 8 MZUSP 36107 7 1 6 Espirito Santo 2446841283 2012-07-09 MZUSP J. B. Mendonca Brazil 14 -20.515862 Enseada da Cachoeira 1 -29.337557 Enseada da Cachoeira 7 8 MZUSP 36131 4 1 3 Espirito Santo 2446841314 2012-02-06 MZUSP L. R. Simone Brazil 14 -20.479914 Praia dos Andradas 1 -29.306675 Enseada da Cachoeira 7 8 MZUSP 36100 2 1 1 Espirito Santo 2446841363 2013-07-17 SCUBA MZUSP J. B. Mendonca Brazil 10 -20.479914 Praia dos Andradas 1 -29.30686 Enseada da Cachoeira 7 8 MZUSP 36089 3 2 1 Espirito Santo 2446841498 2013-07-05 MZUSP J. B. Mendonca Brazil 10 -20.515028 Praia das Tartarugas 1 -29.305056 Enseada da Cachoeira 7 8 MZUSP 36166 3 2 1 Espirito Santo 2446841480 2012-06-26 MZUSP J. B. Mendonca Brazil 10 -20.517027 Parcel 1 -29.299139 Enseada da Cachoeira 7 8 MZUSP 36096 2 2 Espirito Santo 2446841471 2016-06-19 MZUSP Parcel das Tartarugas & J. B. Mendonca Brazil 10 -20.519556 Parcel 1 -29.299555 Enseada da Cachoeira 7 8 MZUSP 36111 2 2 Espirito Santo 2446840357 2019-02-02 MZUSP J. B. Mendonca Brazil 5 -20.519556 Parcel 1 -29.299555 Enseada da Cachoeira 7 8 MZUSP 39876 1 1 Espirito Santo 2446840771 2017-11-15 MZUSP J. B. Mendonca Brazil 5 -20.519556 Parcel 1 -29.299555 Enseada da Cachoeira 7 8 MZUSP 39877 1 1 Espirito Santo 2446841463 2012-07-15 MZUSP J. B. Mendonca Brazil 5 -20.519833 Praia do Parcel 1 -29.300165 Enseada da Cachoeira 7 8 MZUSP 36164 17 4 11 2 Espirito Santo 2446840244 2012-06-19 MZUSP J. B. Mendonca Sao Tome and Principe 12 -20.52225 Praia 1 -29.309639 Enseada da Cachoeira 7 8 MZUSP 36091 4 3 1 Espirito Santo 2446841165 2013-07-16 MZUSP J. B. Mendonca Sao Tome and Principe 10 -20.526527 Pedra da Garoupa 1 -29.309444 Enseada 7 8 MZUSP 36117 4 1 3 Espirito Santo 2446840801 2013-07-09 MZUSP J. B. Mendonca Sao Tome and Principe 10 -20.522888 Farrilhoes 1 -29.331112 Enseada da Cachoeira 7 8 MZUSP 36133 13 5 8 Espirito Santo 2446840672 2012-06-20 MZUSP J. B. Mendonca Sao Tome and Principe 12 -20.522888 Farrilhoes 1 -29.331112 Enseada da Cachoeira 7 8 MZUSP 36109 23 7 16 Espirito Santo 2446841014 2014-10-25 MZUSP J. B. Mendonca Sao Tome and Principe 11 -20.522888 Farrilhoes 1 -29.331112 Enseada da Cachoeira 7 8 MZUSP 36136 2 1 1 Espirito Santo 2446841381 2014-04-22 MZUSP J. B. Mendonca Sao Tome and Principe 15 -20.524944 Paredao 1 -29.328861 Praia do Lixo 7 8 MZUSP 36162 2 1 1 Espirito Santo 2446840587 2012-07-02 MZUSP J. B. Mendonca Sao Tome and Principe 25 -20.524944 Enseada do Lixo 1 -29.328861 Praia do Lixo 7 8 MZUSP 36116 1 1 Espirito Santo 2446840800 2012-02-17 MZUSP C. H. Guimaraes Sao Tome and Principe 23 -20.52875 Enseada do Lixo 1 -29.324472 Praia do Lixo 7 8 MZUSP 36075 1 1 Espirito Santo 2446840896 2012-10-13 MZUSP L. R. L Simone Barbados 13.075213 Turtle Beach 1 -59.604687 7 8 MZUSP 28061 10 4 1 5 Christ Church 2446841435 [264,1415,1988,2016] L. R. L. Simone Grenada 12.029355 Grand Anse Bay 1 -61.771854 7 8 1 1 2446841067 2013-07-13 G. Guzman Costa Rica 5 9.657778 Puerto Viejo 21 -82.755554 8 9 6 6 2446841018 1999-07-22 MZUSP Souza Brazil Fernando de Noronha Simone Praia do Meio 8 9 MZUSP 13894 1 1 Pernambuco 2446841334 1965-09 MZUSP Abrolhos & W. Narchi Brazil Bahia 8 9 MZUSP 17712 1 1 Bahia 2446840795 1985-01-21 MZUSP P. Young Brazil Ilha Redonda Abrolhos 8 9 MZUSP 7028 4 4 Bahia 2446841507 2011-06-15 MZUSP Anker & Feitosa Brazil Meireles Fortaleza 8 9 MZUSP 34990 1 1 Ceara 2465041975 2010-03-28 MZUSP A. Jorio Brazil 22 9 10 -20.84939 Piuma 1 -40.473694 Guarapari 8 9 MZUSP 29139 1 2 Espirito Santo 2465041971 2008-07-18 MZUSP R. C. Marques Brazil 22 Praia do Ermitao Guarapari 9 10 MZUSP 33644 2 2 Espirito Santo 2446841242 2001-01-05 MZUSP de Guanabara Brazil Forte de Santa Cruz 9 10 MZUSP 36746 1 Rio de Janeiro 2446840846 1987-01-10 MZUSP W. Vergara Brazil Praia de Itaipu 9 10 MZUSP 8666 2 2 Rio de Janeiro 2465041974 2015-02-26 2015-02-27 2015-02-26 MZUSP E. P. Goncalves Brazil -23.816668 Sao Sebastiao 1254 -45.383335 Ilha Bela 9 10 MZUSP 12991 3 3 Rio de Janeiro 2465041972 2015-02-26 2015-02-27 2015-02-26 MZUSP E. P. Goncalves Brazil 2 -23.816668 Sao Sebastiao 1254 -45.383335 Ilha Bela 9 10 MZUSP 12991 3 3 Rio de Janeiro 2465041973 1986-06-16 MZUSP W. L. Vergara Brazil 2 Praia da Armacao Santa Catarina 9 10 MZUSP 9431 5 5 2446841070 2011-11-03 MZUSP Santa Catarina Brazil Brazil 9 10 MZUSP 35066 2 2