Beetles that live with ants (Coleoptera, Carabidae, Pseudomorphini): A remarkable new genus and species from Guyane (French Guiana), Guyanemorpha spectabilis gen. n., sp. n. Erwin, Terry L. ZooKeys 2013 2013-12-03 358 11 23 http://zoobank.org/66A2E5B7-0831-4E1E-8AD2-ECE683F0AD8D Erwin Erwin 2013 Hexapoda Carabidae Guyanemorpha GBIF Animalia Guyanemorpha Coleoptera 2 13 Arthropoda genus gen. n.  Type species.   Guyanemorpha spectabilisErwin, sp. n.  Proposed english vernacular generic name. Guyane False-form beetles  Adelphotaxon. Probably  Notopseudomorpha(Baehr, 1997) (see Erwin and Geraci 2008for phylogeny).  Description.  Head( Fig. 2) without supraorbital setigerous punctures, nor any accessory setae; frontal impressions absent. Labrum barely visible with anterior margin shallowly emarginate, quadrisetose; clypeus markedly wide, nearly obscured in dorsal aspect by protruding frons, with obtuse setiferous lateral corners. Eyes slightly convex; small gena with numerous stout setae. Antenna short, just reaching anterior coxa in repose; antennomeres 3-9 slightly wider than 1-3, and appearing slightly flattened. Mandible markedly flattened with a very short and acute apex; outer margin ventral of the scrobe without short stoutsetae. Maxillary palpi markedly short, 3-segmented, palpomeres slightly depressed, palpomere 3 truncate apically. Labial palpus with short bisetose palpomere 2; palpomere 3 markedly securiform and robust, its distal margin mostly membranous with sensory organs.   Figure 2-3. 2  Guyanemorpha spectabilissp. n., female holotype, ADP132101; Risquetout, PK20, Guyane. Head, dorsal planar aspect 3  Guyanemorpha spectabilissp. n., female paratype, ADP132105; Nouragues, Saut Parare, Guyane. Left flight wing, dorsal aspect.  Prothorax.Pronotum ( Fig. 1) much wider than head, transverse, with narrowly beaded margins; without a pair of setigerous punctures each side, apical, lateral and posterior margins with fringe of short stout setae; hind angles obtuse, broadly rounded. Prosternal process unisetiferous subapically, intercoxal process feebly beaded.  Pterothorax.Metepisternum elongate and markedly narrowed posteriorly, the outer margin about 2 times greater in length than the anterior margin, posterior margin about 0.2 times anterior margin.  Elytra.Elytron ( Fig. 1) tapered, markedly narrower apically, width of elytra about equal to that of pronotum at widest point, apical margin truncate with evenly rounded humerus, interneurs and intervals effaced; parascutellar stria absent, scutellum hidden, parascutellar puncture present, marked; without fixed discal setae, surface glabrous. Lateral marginal (umbilical) series of 10 setae, arrayed throughout and widely spaced; lateral margin with fringe of short stout setae.  Hind wings.Macropterous. Venation ( Fig. 3).  Legs.Short and depressed, femur posteriorly channeled for reception of tibia in repose; antennal comb notch very shallow; tibial spurs normal; anterior tarsi of male with tarsomeres 2-4 dilated slightly, ventrally each with two laterally placed rows of adhesive articulo-setae.  Abdomen.Abdominal sterna III-VII with patches of short setae and each of IV-VII with a single row of erect ambulatory setae numbering 2 to 8 setae; V and VI in male each with dense row of yellowish robust setae separated medially.  Male genitalia( Fig. 4). Phallobase hooded with small orifice, dorsum not crested; phalloshaft arched throughout its length, diameter sub-rounded to somewhat depressed dorso-ventrally; phalloapex produced, sharp, rounded, markedly depressed dorso-ventrally; endophallus orifice elongate, endophallus with dense patches of microtrichia. Parameres (C) moderately short compared to those of genus  Pseudomorpha, nearly equal in length, left slightly longer and much broader than right, each apically glabrous. Ring sclerite (E) normal for family.   Figure 4.  Guyanemorpha spectabilissp. n., male paratype, ADP132103; Risquetout, PK20, Guyane. Male genitalia, median lobe and parameres labeled as in repose in male Aventral aspect Bright lateral aspect Cleft lateral aspect Ddorsal aspect.  Female ovipositor and genital tract( Fig. 1). Gonocoxite 2 falcate, base about as long as blade, latter relatively short, pointed distally; margins without ensiform setae; with short preapical nematiform seta.