Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae) Vandenberg, Natalia J. Hanson, Paul E. Zootaxa 2019 2019-02-08 4554 1 255 285 Gordon 1999 [451,552,692,718] Insecta Coccinellidae GBIF Animalia Coleoptera 7 262 Arthropoda tribe Diomini    1. Antenna with 10 antennomeres........................................................................... 2  1’. Antenna with 11 antennomeres........................................................................... 4   2. Species restricted to the New World....................................................................... 3  2’. Species occurring outside the New World............................................... DiomusMulsant, in part   3. Female genitalia ( Fig. 5) with sperm duct generally long, convoluted; spermatheca vermiform throughout, lacking bulbous basal chamber. Male genitalia ( Fig. 7) with main tube of penis completing half to three-quarters turn or more within basal ½, somewhat spiraliform; penis bearing long or short apical flagellum; penis capsule ( Figs. 7, 11–12) with long slender inner arm that may be strongly procurved ( Fig. 12); outer arm truncated or obliterated ( Fig. 11), leaving large oval orifice to mark its place; some species with fragment of outer arm present but not enclosing ejaculatory duct ( Fig. 12). Predators on Sternorrhyncha( Pseudococcidae, Coccidae, Diaspididae, Monophlebidae, Ortheziidae). Larvae free-living.........  DecadiomusChapin  3’. Female genitalia ( Fig. 6) with sperm duct of moderate length, may be weakly sinuous; spermatheca with bulbous basal chamber. Male genitalia ( Figs. 8, 29–32) with main tube of penis completing half turn within basal ½, forming fishhook shape; penis lacking apical flagellum; inner and outer arms of penis capsule ( Fig. 10) equally well developed, broadly consolidated into fanlike shape with sinuous distal margin; ejaculatory duct enters penis capsule through small opening in outer angle of capsule. Parasites of  Piperplants that feed on food bodies. Larvae develop in silken tents constructed between leaf veins ( Figs. 40–42)..................................................................................  Moiradiomus  gen. nov.   4. Length of terminal antennomere approximately equal to twice width, longer than basal 3 antennomeres combined.................................................................................................  AndrzejŚlipiński  4.’ Length of terminal antennomere ( Figs. 1a,b) less than twice width (except in  Diomus sedani), much shorter than basal 3 antennomeres combined..................................................................................... 5   5. Pronotum with sharp submarginal carinae separating the anterolateral pronotal angles from the disc. Australia.....................................................................................................  DichainaWeise  5.’ Pronotum without submarginal carinae..................................................................... 6   6. Prosternum T-shaped, long anterior to coxal cavities (subequal to length of cavity), with short carinae not extending to apex. Female genitalia with bulbous base to spermatheca, with distally tapered sclerotized sheath in basal half of sperm duct (similar to  Moiradiomus; Fig. 6). Restricted to the New World........................................  HeterodiomusBrèthes  6.’ Without the above combination of characters. Distributed world wide................................ DiomusMulsant