Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini) Fominykh, Dmitriy D. Zamotajlov, Alexandr S. Khomitskiy, Eugeniy E. Titarenko, Andrey Yu. Zootaxa 2020 2020-01-30 4729 3 301 328 8M7WN Fominykh & Zamotajlov & Khomitskiy & Titarenko, 2020 Fominykh & Zamotajlov & Khomitskiy & Titarenko 2020 [181,693,153,180] Insecta Carabidae Carabus Animalia Coleoptera 4 305 Arthropoda subSpecies stjernvalli subsp. nov. Lipaster paravanicus   urn:lsid:zoobank.org:act: 7F961A11-E05D-48CD-8BCD-36E08FE17D4E ( Figs. 1, 3–6, 92, 115–118)   Type material.  Holotype: ♂, Georgia, Samtskhe-Dzhavakheti, Wbank of Paravani Lakenear Asparavill., E slopes of Kyerogly Mt.,  2100-2200 m, N 41°28’24.99”/ E 43°46’19.65”, 04.-  05.06.2015( D. Fominykh, A. Zubov& A. Safronov) ( ZISP).  Paratypes: 444♂♂, 490♀♀, same data as holotype, pitfall traps( ZISP, MPSU, CFDD, CTAY, CZAM, CKEE, CSAA).   Description.Medium sized subspecies ( Figs. 3–6). GBL = 26.8–34.8 mm, males 26.8–30.0 mm, females 28.6– 34.8 mm, SBL = 23.6–31.4 mmin both sexes. Body rather robust, ovate. Underside black, dorsum generally with bright metallic luster, green, bronze, more seldom violet or blue, in females less bright, somewhat mat; pronotum basally and elytra with metallic luster. Head broad, a little narrower than pronotum, HL = 4.0– 4.8 mmin males, 4.2–5.6 mmin females, HW = 4.8–6.0 mm in males, 5.4–6.6 mmin females; antennae short. Pronotum transverse, PW = 6.6–8.6 mmin males, 7.8–9.8 mmin females, PB = 5.4–6.8 mmin males, 6.0– 7.8 mmin females, PL = 4.0–5.0 mm in males, 4.4–5.6 mmin females, PW/HW = 1.2–1.6 inboth sexes, PW/PL = 1.2–1.6 inboth sexes, PW/PB = 1.1–1.4 inboth sexes; hind angles faintly protruding posteriorly; disk rugose-punctate. Legs short. Elytra convex, EW/PW = 1.2–1.5 inboth sexes; EL = 15.6–17.8 mmin males, 17.2–20.8 mmin females, EW = 9.8–11.4 mmin males, 10.4–12.6 mmin females, EL/EW = 1.5–1.8 inboth sexes; elytral punctation faint and sparse, usually with only several rarefied impressions behind. Male genitalia ( Fig. 92) in general shape of the projections and lobes of endophallus resemble other forms of  C. stjernvalli.   Differential diagnosis and remarks.This subspecies differs from the other  C. stjernvallitaxa, first of all, in its body proportions ( Table 1, Fig. 116). In particular, it is distinguishable from  C. stjernvalli stjernvalliand  C. stjernvalli gvalijaiby broader elytra. Among all known forms it is the closest morphologically to  C. stjernvalli tabackuriensis, but is distinguishable by body size and some proportions. The new subspecies also differs from all other taxa of  C. stjernvalliin the structure of endophallus ( Fig. 92), particularly ventroapical and ventrobasal lobes are more pronounced, also a small projection, missing in the other populations from Trialeti Mt. Range and Dzhavakheti Upland, is present beneath ventrobasal lobe.   Distribution and habitat.The new subspecies populates west spurs of the Samsar Mt. Range at western bank of Paravani Lake. It inhabits alpine meadows at 2100–2200 m. The imago are active from the beginning of June. The following  Carabusspecies occur together with this subspecies:  C. armeniacusMannerheim, 1830,  C. calleyiFischer von Waldheim, 1823,  C. cribratusQuensel, 1806, and  C. maurusAdams, 1817.   Etymology.The subspecific epithet refers to the typelocality, vicinity of Paravani Lake in Georgia. 2561484965 2015-05-06 ZISP Samtskhe-Dzhavakheti, W & D. Fominykh & A. Zubov & A. Safronov Georgia Kyerogly Mt. 2150 41.47361 Paravani Lake 1 43.772125 Aspara 4 305 1 1 holotype 2561484959 2015-05-06 ZISP, MPSU, CFDD, CTAY, CZAM, CKEE, CSAA Samtskhe-Dzhavakheti, W & D. Fominykh & A. Zubov & A. Safronov Georgia Kyerogly Mt. 2150 41.47361 Paravani Lake 1 43.772125 Aspara 4 305 934 490 444 paratype