New species in Rejectaria Guenée (Lepidoptera: Erebidae: Herminiinae) with a focus on the Cyclanthaceae-feeders Goldstein, Paul Z. Janzen, Daniel H. Hallwachs, Winnie Proshek, Benjamin Zootaxa 2022 2022-01-07 5087 3 451 483 Goldstein Goldstein & Janzen & Hallwachs & Proshek 2022 [151,586,1840,1867] Insecta Noctuidae Rejectaria Animalia Lepidoptera 16 467 Arthropoda species richardashleyi sp. nov. Goldstein & Janzen & Hallwachs & Proshek 2022 [151,252,1479,1504] Insecta Noctuidae Rejectaria Animalia Lepidoptera 25 476 Arthropoda species paratrax  Habitus Figs 21–22, 35–36, 42. Male genitalia Figs 56, 83–84. Female genitalia Figs 94, 102. Larva Figs 115–117.   Material examined. Type material:   Holotype ♂  COSTA RICA: Guanacaste, Area de Conservacion Guanacaste, Sector Pitilla: Sendero Mismo, 10.98758, -85.41967, el.  680m, larva on  Asplundia microphylla,  09/16/2009, ecl.  10/20/2009, Jose Calvo, collector, 09-SRNP-32559, USNMENT01493269.   Paratypes( 68♂♂, 56♀♀)   COSTA RICA: ALAJUELA, AREADE CONSERVACION GUANACASTE. SECTOR RINCON RAIN FOREST( 33♂♂, 28♀♀): 1♂, Comejen, 10.95669, -85.28934, el.  213m, larva on  Asplundia microphylla,  11/05/2012, ecl.  12/04/2012, Keiner Aragon, collector, 12-SRNP-68765, USNMENT01493114.  1♂, Laureles, 10.93319, -85.25335, el.  95m, larva on  Asplundia utilis,  06/10/2007, Jose Perez, collector, ecl.  07/10/2007, 07- SRNP-41583, USNMENT01493209.  1♂, 1♀ Palomo, 10.96187, -85.28045, el.  96m, larva on  Asplundia utilis,  12/14/2010, Manuel Rios, collector: ( 1♂) ecl.  01/13/2011, 10-SRNP-68137, USNMENT01493014; ( 1♀) ecl.  01/12/2011, 10-SRNP-68139, USNMENT01493079.  5♂♂, 6♀♀, Quebrada Escondida, 10.89928, -85.27486, el.  420m, larva on  Asplundia utilis: ( 1♂)  10/09/2001, ecl.  11/09/2001, Freyci Vargas, collector, 01-SRNP-5975, USNMENT01493279; ( 1♂)  12/05/2001, ecl.  12/30/2001, Freyci Vargas, collector, 01-SRNP-23447, USNMENT01479307; ( 1♂)  12/18/2001, ecl.  01/24/2002, Jose Perez, collector, 01-SRNP-23524, USNMENT01493009; ( 1♂)  11/29/2010, ecl.  12/29/2010, Anabelle Cordoba, collector, 10-SRNP-44494, USNMENT01493153; ( 1♂)  02/25/2011, ecl.  03/29/2011, Jose Perez, collector, 11-SRNP-41004, USNMENT01493262. ( 1♀)  10/09/2001, ecl.  11/10/2001, Freyci Vargas, collector, 01-SRNP-5973, USNMENT01493138; ( 1♀)  12/05/2001, ecl.  12/28/2001, Freyci Vargas, collector, 01-SRNP-23446, USNMENT01493117; ( 1♀)  09/17/2003, ecl.  10/09/2003, Jose Perez, collector, 03-SRNP-12755.1, USNMENT01493141; ( 1♀)  08/06/2005, ecl.  08/29/2005, Jose Perez, collector, 05-SRNP-42034, USNMENT01493167; ( 1♀)  11/25/2009, ecl.  12/16/2009, Jose Perez, collector, 09-SRNP-43306, USNMENT01493290; ( 1♀)  02/25/2011, ecl.  03/28/2011, Jose Perez, collector, 11-SRNP-41005, USNMENT01493033.  1♂, 1♀, Quebrada Escondida, 10.89928, -85.27486, el.  420m, larva on  Carludovica costaricensis,  12/05/2013, Jose Perez, collector: ( 1♂) ecl.  01/05/2014, 13-SRNP-44863, USNMENT01493237; ( 1♀) ecl.  01/01/2014, 13-SRNP-44864, USNMENT01493160.  3♂♂, 2♀♀, Quebrada Guarumo, 10.90445, - 85.28412, el.  400m, larva on  Asplundia utilis,Jose Perez, collector,  09/28/2010: ( 1♂) ecl.  10/24/2010, 10-SRNP- 43538, USNMENT01493186; ( 1♂) ecl.  10/24/2010, 10-SRNP-43540, USNMENT01493206; ( 1♂) ecl.  10/24/2010, 10-SRNP-43541, USNMENT01493204; ( 1♀) ecl.  10/18/2010, 10-SRNP-43539, USNMENT01493245; ( 1♀) ecl.  10/31/2010r, 10-SRNP-43542, USNMENT01493005. 2♂♂, 1♀, Rio Francia Arriba, 10.89666, -85.29003, el.  400m: larva on  Asplundia utilis: ( 1♂)  10/07/2008, ecl.  11/01/2008, Anabelle Cordoba, collector, 08-SRNP-41982, USNMENT01493082; ( 1♂)  07/08/2013, Jose Perez, collector, ecl.  08/04/2013, 13-SRNP-42618, USNMENT01493092; ( 1♀) same data as previous, ecl.  08/07/2013, 13-SRNP-42617, USNMENT01493172.  1♂, Selva, 10.92291, -85.31877, el.  410m, Larva on  Asplundia utilis,  12/16/2016, ecl.  01/08/2017, Edwin Apu, collector, 16-SRNP-81079, USNMENT01501726.  2♂♂, 4♀♀ Sendero Anonas, 10.90528, -85.27882, el.  405m, larva on  Asplundia utilis ( 1♂) Jose Perez, collector,  10/10/2003, ecl.  11/09/2003, 03-SRNP-31011, USNMENT01493059; ( 1♀)  08/09/2004, ecl.  09/13/2004, Jose Perez, collector, 04-SRNP-42008, USNMENT01493062; ( 1♂)  02/27/2013, ecl.  04/07/2013, 13-SRNP-40884, USNMENT01493179; ( 1♀)  02/27/2013, ecl.  03/28/2013, 13-SRNP-40883, USNMENT01493162; ( 1♀)  12/02/2014, ecl.  12/18/2014, Anabelle Cordoba, collector, 14-SRNP-44987, USNMENT01493244; ( 1♀)  09/19/2005, ecl.  10/18/2005, Minor Carmona, collector, 05-SRNP-42715, USNMENT01493022.  2♀♀, Sendero Anonas, 10.90528, -85.27882, el.  405m, larva on  Carludovica costaricensis  02/12/2013, Pablo Umaña Calderon, collector: ( 1♀) ecl.  03/17/2013, 13-SRNP-40664, USNMENT01493278; ( 1♀) ecl.  03/13/2013, 13-SRNP-40665, USNMENT01493154.  1♀, Jacobo, 10.94076, -85.3177, el.  461m, larva on  Asplundia utilis,  12/02/2010, ecl.  01/03/2011, Manuel Ríos, collector, 10-SRNP-81463, USNMENT01493115.  2♀♀, Sendero Guaca, 10.9061, -85.28281, el.  400m, ( 1♀) larva on  Cyclanthus bipartitus,  12/27/2002, ecl.  01/24/2003, Jose Perez, collector, 02-SRNP-21559, USNMENT01493222; ( 1♀) larva on  Asplundia utilis,  12/11/2003, ecl.  01/09/2004, Jose Perez, collector, 03-SRNP-31713, USNMENT01493037.  10♂♂, 4♀♀, Sendero Rincon, 10.8962, -85.27769, el.  430m, larva on  Asplundia utilis: ( 1♂)  01/09/2002, ecl.  02/10/2002, Armando Rios, collector, 02-SRNP-6051, USNMENT01493250; ( 1♂)  02/12/2002, ecl.  03/15/2002, Jose Perez, collector, 02- SRNP-6319, USNMENT01493282; ( 1♂)  11/13/2004, ecl.  12/13/2004, Jose Perez, collector, 04-SRNP-42741, USNMENT01493036; ( 1♂)  11/25/2004, ecl.  12/27/2004, Jose Perez, collector, 04-SRNP-42810, USNMENT01493150; ( 1♂)  12/09/2004, ecl.  01/09/2005, Jose Perez, collector, 04-SRNP-42900, USNMENT01493187, USNMslide 148578; ( 1♂)  01/02/2005, ecl.  02/01/2005, Jose Perez, collector, 05-SRNP- 40001, USNMENT01493118; ( 1♂)  01/20/2005, ecl.  02/28/2005, Minor Carmona, collector, 05-SRNP-40228, USNMENT01493241; ( 1♂)  12/07/2005, ecl.  01/09/2006, Jose Perez, collector, 05-SRNP-43616, USNMENT01493078; ( 1♂)  11/07/2008, ecl.  12/08/2008, Jose Perez, collector, 08-SRNP-42148, USNMENT01493083; ( 1♂)  11/07/2008, ecl.  12/11/2008, Jose Perez, collector, 08-SRNP-42149, USNMENT01493042;( 1♀)  01/09/2002,ecl.  02/06/2002, JosePerez,collector,02-SRNP-6048,USNMENT01493284; ( 1♀)  01/09/2002, ecl.  02/06/2002, Armando Rios, collector, 02-SRNP-6053, USNMENT01493252; ( 1♀)  11/07/2008, ecl.  12/09/2008, Jose Perez, collector, 08-SRNP-42147, USNMENT01493286; ( 1♀)  02/26/2013, ecl.  03/24/2013, Jose Perez, collector, 13-SRNP-40837, USNMENT01493292.  1♀, Sendero Rincon, 10.8962, - 85.27769, el.  430m, larva on  Cyclanthus bipartitus,  09/17/2010, ecl.  10/07/2010, Pablo Umaña Calderon, collector, 10-SRNP-43308, USNMENT01493077.  2♂♂, 2♀♀, Sendero Venado, 10.89678, -85.27001, el.  420m, larva on  Asplundia utilis, Jose Perez, collector: ( 1♂)  12/03/2004, ecl.  01/10/2005, 04-SRNP-42863, USNMENT01493101; ( 1♂)  10/23/2013, ecl.  11/29/2013, 13-SRNP-44157, USNMENT01493164; ( 1♀)  12/03/2004, ecl.  01/08/2005, 04- SRNP-42864, USNMENT01493227; ( 1♀)  10/23/2013, ecl.  11/27/2013, 13-SRNP-44156, USNMENT01493263.  1♂, Sendero Juntas, 10.90661, -85.28784, el.  400m, larva on  Asplundia utilis,  11/17/2010, ecl.  12/20/2010, Jose Perez, collector, 10-SRNP-44288, USNMENT01479216.  2♂♂, 1♀, Vado Rio Francia, 10.90093, -85.28915, el.  400m, Jose Perez, collector, larva on  Asplundia utilis: ( 1♂)  01/07/2002, ecl.  02/09/2002, 02-SRNP-6038, USNMENT01493148; ( 1♂)  01/02/2015, ecl.  01/31/2015, 15-SRNP-40024, USNMENT01479359; ( 1♀)  11/01/2013, ecl.  12/06/2013, 13-SRNP-44276, USNMENT01493094.  1♂, Vado Rio Francia, 10.90093, -85.28915, el.  400m, Jose Perez, collector, larva on  Carludovica costaricensis,  01/11/2013, ecl.  02/15/2013, 13-SRNP-40115, USNMENT01493132.  SECTOR SAN CRISTOBAL( 9♂♂, 8♀♀): 3♂♂, Rio Blanco Abajo, 10.90037, -85.37254, el.  500m: ( 1♂)  11/15/2004, ecl.  12/09/2004, larva on  Asplundia utilis, Gloria Sihezar, collector, 04-SRNP-60897, USNMENT01493012; ( 1♂)  11/05/2005, ecl.  12/08/2005, larva on  Asplundia utilis, Gloria Sihezar, collector, 05- SRNP-6913, USNMENT01493066; ( 1♂) larva on  Asplundia microphylla,  12/04/2005, ecl.  12/27/2005, Osvaldo Espinoza, collector, 05-SRNP-7574, USNMENT01493091.  2♂♂, 1♀, Sendero Colegio, 10.89296, -85.3788, el.  520m, Larva on  Asplundia utilis, Gloria Sihezar, collector: ( 1♂)  10/03/2005, ecl.  11/09/2005, 05-SRNP-6176, USNMENT01493120; ( 1♀)  10/11/2006, ecl.  11/11/2006, 06-SRNP-8361, USNMENT01493229; ( 1♂) larva on  Cyclanthus bipartitus,  08/25/2006, ecl.  09/18/2006, Carolina Cano, collector,06-SRNP-7082, USNMENT01493144.  1♂, 1♀, Sendero Huerta, 10.9305, -85.37223, el.  527m, larva on  Asplundia utilis,  01/16/2007, Osvaldo Espinoza, collector: ( 1♂) ecl.  02/16/2007, 07-SRNP-273, USNMENT01493063; ( 1♀) ecl.  02/17/2007, 07-SRNP-271, USNMENT01493289.  1♀, Brasilia: Moga, 11.01227, -85.34929, el.  320m, larva on  Asplundia utilis,  12/03/2013, ecl.  12/30/2013, Minor Carmona, collector, 13-SRNP-65705, USNMENT01493048.  2♂♂, 2♀♀, Sendero Palo Alto, 10.88186, -85.38221, el.  570m, larva on  Asplundia utilis: ( 1♂)  01/16/2017, Elda Araya, collector, ecl.  02/10/2017, 17-SRNP-67, USNMENT01501706; ( 1♂)  01/16/2017, Elda Araya, collector, ecl.  02/13/201717- SRNP-69, USNMENT01501703; ( 1♀)  09/20/2005, Carolina Cano, collector, ecl.  10/16/2005, 05-SRNP-5882, USNMENT01493194; ( 1♀)  09/20/2005, Carolina Cano, collector, ecl.  10/18/2005, 05-SRNP-5883, USNMENT01493264.  1♂, 3♀♀, Tajo Angeles, 10.86472, -85.41531, el.  540m, larva on  Asplundia utilis, Gloria Sihezar, collector,  09/20/2010: ( 1♂) ecl.  10/24/2010, 10-SRNP-5402, USNMENT01493126; ( 1♀) ecl.  10/11/2010, 10-SRNP-5397, USNMENT01493034; ( 1♀) ecl.  10/16/2010, 10-SRNP-5399, USNMENT01493032; ( 1♀) ecl.  10/22/2010, 10-SRNP-5401, USNMENT01493166.  SECTORDEL ORO ( 5♂♂, 1♀): 4♂♂, 1♀, Bosque Aguirre, 11.0006, -85.438, el.  620m, larva on  Cyclanthus bipartitus,  11/11/2010, Lucia Ríos, collector: ( 1♂) ecl.  12/11/2010, 10-SRNP-22482, USNMENT01493142: ( 1♂) ecl.  12/12/2010, 10-SRNP-22479, USNMENT01493035; ( 1♂) ecl.  12/12/2010, 10-SRNP-22480, USNMENT01493193;( 1♂)ecl.  12/20/2010, 10-SRNP-22481, USNMENT01493137; ( 1♀)  05/24/2015, ecl.  06/22/2015, 15-SRNP-20615, USNMENT01493202. 1♂, Monte Cristo, 11.01373, -85.42531, el.  525m, larva on  Cyclanthus bipartitus,  11/07/2016, ecl.  12/03/2016, Lucia Ríos, collector, 16-SRNP-21701, USNMENT01501728.  SECTORPITILLA ( 21♂♂, 16♀♀): 4♂♂, 3♀♀, Sendero Cuestona, 10.99455, -85.41461, el.  640m, larva on  Asplundia microphylla: ( 1♂)  12/06/2003, ecl.  01/08/2004, Lucia Ríos, collector, 03-SRNP-37286, USNMENT01493055; ( 1♀)  10/03/2006, ecl.  10/27/2006, Manuel Rios, collector, 06-SRNP-34630, USNMENT01493192; ( 1♂)  07/01/2011, ecl.  09/01/2011, Manuel Rios, collector, 11-SRNP-31890, USNMENT01493260; ( 1♀)  08/25/2011, ecl.  09/25/2011, Manuel Rios, collector, 11-SRNP-32415, USNMENT01493105; ( 1♂)  09/28/2011, ecl.  11/03/2011, Manuel Rios, collector, 11-SRNP-32933, USNMENT01493113; ( 1♂)  11/15/2012, Calixto Moraga, collector: ecl.  12/26/2012, 12-SRNP-31712, USNMENT01493076; ( 1♀) ecl.  12/20/2012, 12-SRNP-31711, USNMENT01493224.  2♂♂, Sendero Evangelista, 10.9868, -85.42083, el.  660m: larva on  Asplundia microphylla: ( 1♂)  08/03/2011: ecl.  09/01/2011, Freddy Quesada, collector, 11-SRNP-32123, USNMENT01493257; ( 1♂)  08/05/2013, ecl.  09/04/2013, Manuel Rios, collector, 13- SRNP-31037, USNMENT01493276.  7♂♂, 4♀♀, Sendero Laguna, 10.9888, -85.42336, el.  680m, larva on  Asplundia microphylla: ( 1♂)  08/25/2004, ecl.  09/20/2004, Calixto Moraga, collector, 04-SRNP-34808, USNMENT01493228; ( 1♂)  10/07/2006, ecl.  11/06/2006, Manuel Rios, collector, 06-SRNP-34667, USNMENT01493028; ( 1♂)  12/22/2008, ecl.  01/26/2009, Manuel Rios, collector, 08-SRNP-33140, USNMENT01493088 ( 1♂)  11/23/2009, ecl.  12/25/2009, Calixto Moraga, collector, 09-SRNP-33319, USNMENT01493149; ( 1♂)  08/15/2011, ecl.  09/07/2011, Freddy Quesada, collector, 11-SRNP-32330, USNMENT01493235; ( 1♂)  02/19/2013, ecl.  04/06/2013, Calixto Moraga, collector, 13-SRNP-30344, USNMENT01493016; ( 1♂)  08/26/2013, ecl.  10/02/2013, Freddy Quesada, collector, 13-SRNP-31215, USNMENT01493047; ( 1♀)  10/09/2006, ecl.  11/02/2006, Petrona Rios, collector, 06-SRNP- 34701, USNMENT01493089; ( 1♀)  12/09/2008, ecl.  01/11/2009, Petrona Rios, collector, 08-SRNP-33069, USNMENT01493002; ( 1♀)  12/09/2008, ecl.  01/15/2009, Petrona Rios, collector, 08-SRNP-33070, USNMENT01493163; ( 1♀)  12/22/2014, ecl.  01/25/2015, Freddy Quesada, collector, 14-SRNP-31775, USNMENT01493080.  1♂, 1♀, Sendero Memos, 10.98171, -85.42785, el.  740m, larva on  Asplundia microphylla: ( 1♂)  11/28/2003, ecl.  01/07/2004, Lucia Ríos, collector, 03-SRNP-37132, USNMENT01493211; ( 1♀)  12/04/2007, ecl.  12/30/2007, Manuel Rios, collector, 07-SRNP-34060, USNMENT01493060, USNMslide 148389.  4♂♂, 2♀♀, Sendero Mismo, 10.98758, -85.41967, el.  680m, larva on  Asplundia microphylla: ( 1♂)  09/28/2011, ecl.  11/09/2011, Manuel Rios, collector, 11-SRNP-32934, USNMENT01493253; ( 1♂)  08/22/2013, ecl.  09/24/2013, Freddy Quesada, collector, 13-SRNP-31144, USNMENT01493129; ( 1♂)  08/22/2013, ecl.  10/06/2013, Freddy Quesada, collector, 13-SRNP-31213, USNMENT01493190; ( 1♂)  08/22/2013, ecl.  10/07/2013, Freddy Quesada, collector, 13-SRNP- 31235, USNMENT01493045; ( 1♀)  08/20/2010, ecl.  09/20/2010, Petrona Rios, collector, 10-SRNP-31827, USNMENT01493216,  USNMslide 148577; ( 1♀)  09/14/2016, ecl.  10/12/2016, Freddy Quesada, collector, 16- SRNP-31531, USNMENT01501768.  1♂, Sendero Nacho, 10.98445, -85.42481, el.  710m: larva on  Asplundia microphylla,  09/27/2011, ecl.  10/23/2011, Manuel Rios, collector, 11-SRNP-32907, USNMENT01493104.  2♂♂, 3♀♀, Sendero Naciente, 10.98705, -85.42816, el.  700m, larva on  Asplundia microphylla: ( 1♂)  11/25/2013, ecl.  01/07/2014, Calixto Moraga, collector, 13-SRNP-31690, USNMENT01493214; ( 1♂)  11/25/2013, ecl.  01/04/2014, Calixto Moraga, collector, 13-SRNP-31693, USNMENT01493170; ( 1♀)  10/19/2012, ecl.  11/27/2012, Manuel Rios, collector, 12-SRNP-31546, USNMENT01493130; ( 1♀)  11/19/2014, ecl.  01/02/2015, Freddy Quesada, collector, 14-SRNP-31689, USNMENT01493003; ( 1♀)  11/09/2016, ecl.  12/08/2016, Freddy Quesada, collector, 16-SRNP-31843, USNMENT01501778.  1♀, Sendero Tierra Blanca, 10.98122, -85.41773, el.  680m: larva on  Asplundia microphylla:  03/07/2013,ecl.  04/09/2013, Manuel Rios,collector,13-SRNP-30378,USNMENT01493169.  1♀, Estacion Pitilla, 10.98931, -85.42581, el.  675m: larva on  Asplundia microphylla:  10/02/2014, ecl.  11/06/2014, Freddy Quesada, collector, 14-SRNP-31297, USNMENT01493086.  1♀, Sendero Bernales, 10.9835, -85.42117, el.  660m: larva on  Asplundia microphylla:  01/09/2017, ecl.  02/09/2017, Freddy Quesada, collector, 17-SRNP-30083, USNMENT01501776.   Diagnosis.  Rejectaria richardashleyibears a number of strong similarities to both  R. magas(see diagnosis above) and  B. albidiscalis.It differs from both species in its large hind-femoral tufts; from  albidiscalisin the broken HW submarginal band, which is more continuous in  albidiscalis, and possibly in the configuration of the male palp, the second segment of which extends approximately to just beyond the patagium in  richardashleyi, and to the center of the thorax in  albidiscalis.  Rejectaria richardashleyiis also similar to  Strathocles magnipilosabut the latter species has vastly expanded hind femoral tufts and elongate forewings, each with a posteromedial hyaline patch.  Rejectaria richardashleyiis outwardly distinguished from other Neotropical Herminiinae by the combination of the bold yellowish-tan am and pm lines on the forewing, their dark shading flanking the median field, and the small pale crescent lunule; additionally, in the male genitalia, the costa of the valva is completely fused to the valva, and sclerotized in its basal half; the vesica bears a conspicuously anvil-shaped subbasal lobe, similar to that in  magas. The female genitalia lack the serrate semicircular ridges inside the bursa which are present in  villosa, villavicencia, magas,and  ritaashleyae.   Description. Head. Frons, vertex dark chocolate-brown; antennae setose-ciliate; male palpi ( Fig. 35) dark brown, orange scaling along inner edge of 2 ndsegment, highly developed, sweeping back sharply over the head, but with no acute angles; female palpi ( Fig. 36) cocoa-brown, curved in a semicircle, 3 rdsegment fine, nearly as long as 2 nd, scaled dark gray and faintly edged with white at base and tip.  Thorax. Wings—(males, 19mm, n=5; females, 18mm, n=5) Forewing dusky brown; basal line absent; am and pm lines both present and bicolorous, dark brown outwardly, paler inward; median stigma a small pale lunule; pm line sharp, inwardly a diffuse dark brown band, outwardly a more sharp, tannish brown line, angled slightly inward near costa; st line faint, pale, jagged; terminal line a series of black chevrons edged outwardly with tannish brown; HW concolorous with FW, basal and am lines absent, pm and st lines continuous with those of FW. Legs—( Figs 35, 36) Male with conspicuous tufts on all legs, hind-femoral tufts the most expansive; white bands present in female.   FIGURES 43–50.Male genitalia, valvae. 43  R. villosa, Costa Rica, USNMENT01493304, 13-SRNP-45393, Slide USNM148507. 44 R.sp. nr.  villosa, USNMENT01493015, 02-SRNP-6046, Slide USNM148569. 45  R. villosa, Panama, USNMENT01422998, Slide USNM148665. 46  R. villosa, Venezuela, USNMENT01422978, Slide USNM148658. 47  R. villosa, Bolivia, USNMENT01756824, Slide 148669CMNH. 48  R. villosa, Colombia, USNMENT01422993, Slide USNM148666. 49  R. villavicencia, Costa Rica, USNMENT01493395, 06-SRNP-6834, Slide USNM148571. 50  R. villavicencia, Holotype, Colombia, USNMENT00973787, Slide USNM148642   FIGURES 51–56.Male genitalia, valvae. 51  R. ritaashleyae, Costa Rica, USNMENT01493231, Slide USNM148597. 52  R. paratrax, Panama, USNMENT01756002, Slide USNM148595. 53  R. splendida, Syntype, Costa Rica, USNMENT00973707, Slide USNM148598. 54  Narcaea atrax(=  Rejectaria atrax), Holotype, Ecuador, USNMENT00973692, Slide USNM148640. 55  R. magas, Costa Rica, USNMENT01493483, 04-SRNP-27241, Slide USNM148574. 56  R. richardashleyi, Costa Rica, USNMENT01493187, 04-SRNP-42900, Slide USNM148578   FIGURES 57–64.Male Genitalia, vesicae. 57–58 R._ villosa, Costa Rica, USNMENT01493304, 13-SRNP-45393, Slide USNM148507. 59–60 R.sp. nr.  villosa, Costa Rica, USNMENT01493015, 02-SRNP-6046, Slide USNM148569. 61–62  R. villavicencia, Costa Rica, USNMENT01493395, 06-SRNP-6834, Slide USNM148571. 63–64  R. villavicencia, Holotype, Colombia, USNMENT00973787, Slide USNM148642   FIGURES 65–72.Male Genitalia, vesicae. 65–66 R._ villosa, Panama, USNMENT01422998, Slide USNM148665. 67–68 R._ villosa, Venezuela, USNMENT01422978, Slide USNM148658. 69–70 R._ villosa, Bolivia, USNMENT01756824, Slide USNM148669CMNH. 71–72 R._ villosa, Bolivia, USNMENT01422993, Slide USNM148666   FIGURES 73–80.Male genitalia, vesicae. 73–74  R. ritaashleyae, Costa Rica, USNMENT01493231, 03-SRNP-11914, Slide USNM148597. 75–76  R. splendida, Syntype, Costa Rica, USNMENT00973707, Slide USNM148598. 77–78  Narcaea atrax(=  Rejectaria atrax), Ecuador, Holotype, USNMENT00973692, Slide USNM148640. 79–80  R. paratrax, Panama, USNMENT01756002, Slide USNM148595   FIGURES 81–84.Male genitalia, vesicae. 81–82  R. magas, Costa Rica, USNMENT01493483, 04-SRNP-27241, Slide USNM148574. 83–84  R. richardashleyi, Costa Rica, USNMENT01493187, 04-SRNP-42900, Slide USNM148578  Abdomen. Dark brown, concolorous with uppersides of wings.  Male genitalia. ( Figs 56, 83–84) Uncus elongate, setose, sheepsfoot-shaped, costal margin of valva sclerotized or partially sclerotized basally, without any part free, sclerotized in basal half, a small setal tuft at terminus of sclerotization; vinculum bluntly tapered; phallic ridge present; vesica multi-lobate, with thin sclerotized patch originating on phallus and bearing reduced patch of microspines; microspines on remainder of vesica concentrated primarily on two subbasal lobes, one anvil-shaped.   Immature stages. ( Figs 115–117) Mature larva green with pale subdorsal and spiracular stripes; head green with four dark stripes; dorsal stripes yellowish; subdorsal and spiracular stripes white; intersegmental bands yellowish white.   Etymology.  Rejectaria richardashleyiis named in honor of Richard C.Ashley of Texas and Nicoya, Costa Rica, in recognition of his many years of support for Biodiversity Development and Inventory of Area of Conservacion Guanacasteand its recent expansion nationally as BioAlfa, an effort to support Costa Rica’s bioliteracy.   Biology.Larvae documented feeding on  Asplundia microphylla(n =43),  Asplundia utilis(n =74),  Carludovica costaricensis(n =5), and  Cyclanthus bipartitus(n=9). The parasitoid  Zelomorpha paulgoldsteini( Braconidae: Agathidinae) was reared from  R. richardashleyilarvae on  Asplundia utilis(3 records: 10-SRNP-68136, 10-SRNP- 68138, 15-SRNP-45311); see parasitoid records under  ritaashleyae,above.   Distribution. Costa Rica   Remarks.This species bears a number of similarities to both  Bleptina albdidiscalisand  Strathocles magnipilosaas well as  Rejectaria magas, but in the absence of a phylogenetic analysis it is unclear whether these represent synapomorphies. 3428543379 2009-09-16 2009-10-20 2009-09-16 de Conservacion & Jose Calvo Costa Rica 680 10.98758 Area de Conservacion 1 -85.41967 16 467 1 1 Guanacaste holotype 3428543387 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Keiner Aragon Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543400 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543330 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Manuel Rios Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543382 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Freyci Vargas & Anabelle Cordoba Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543315 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543301 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Anabel Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428568305 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543342 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Anabelle Cordoba Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543357 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543314 [414,1387,1123,1149] 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543397 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Edwin Apu Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543310 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428568301 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428568308 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543359 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Anabelle Cordoba Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543312 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Minor Carmona Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543338 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Pablo Umana Calderon Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543325 [676,1377,1447,1473] 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543347 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543348 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Manuel Rios Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543372 [151,785,1554,1581] 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543340 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543356 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543328 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543332 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Armando Rios Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543374 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543386 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543309 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543390 2001-05-12 2017-08-01 2001-05-12 CONSERVACION Jose Perez Costa Rica Sendero Rincon 213 10.8962 ALAJUELA 5 -85.27769 Comejen 17 468 124 56 68 Alajuela paratype 3428543335 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 17 468 26 11 15 GUANACASTE paratype 3428543366 2002-06-02 2015-02-01 2002-06-02 USNM Minor Carmona Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 17 468 26 11 15 GUANACASTE paratype 3428543346 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 17 468 26 11 15 GUANACASTE paratype 3428543385 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 17 468 26 11 15 GUANACASTE paratype 3428543304 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 18 469 10.90093 Rio Francia 1 -85.28915 Sendero Venado 17 468 26 11 15 GUANACASTE paratype 3428543403 [151,1367,186,213] 2002-06-02 2015-02-01 2002-06-02 USNM Armando Rios Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428568304 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428543319 2002-06-02 2015-02-01 2002-06-02 USNM Pablo Umana Calderon Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428543324 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428568303 2002-06-02 2015-02-01 2002-06-02 USNM Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428543362 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428543344 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428568302 2002-06-02 2015-02-01 2002-06-02 USNM Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428543317 2002-06-02 2015-02-01 2002-06-02 USNM Jose Perez Costa Rica Sendero Rincon 430 10.90093 Rio Francia 1 -85.28915 Sendero Venado 18 469 26 11 15 GUANACASTE paratype 3428568306 2004-09-12 2017-10-02 2004-09-12 SECTOR Gloria Sihezar Saint Kitts and Nevis 500 10.86472 Rio Blanco Abajo 1 -85.41531 18 469 50 23 27 GUANACASTE paratype 3428543401 2004-09-12 2017-10-02 2004-09-12 SECTOR Osvaldo Espinoza Saint Kitts and Nevis 500 10.86472 Rio Blanco Abajo 1 -85.41531 18 469 50 23 27 GUANACASTE paratype 3428543355 2004-09-12 2017-10-02 2004-09-12 SECTOR Gloria Sihezar & Carolina Cano Saint Kitts and Nevis 500 10.86472 Rio Blanco Abajo 1 -85.41531 18 469 50 23 27 GUANACASTE paratype 3428543393 2004-09-12 2017-10-02 2004-09-12 SECTOR Osvaldo Espinoza Saint Kitts and Nevis 500 10.86472 Rio Blanco Abajo 1 -85.41531 18 469 50 23 27 GUANACASTE paratype 3428543363 2004-09-12 2017-10-02 2004-09-12 SECTOR Minor Carmona Saint Kitts and Nevis 500 10.86472 Rio Blanco Abajo 1 -85.41531 18 469 50 23 27 GUANACASTE paratype 3428543389 2004-09-12 2017-10-02 2004-09-12 SECTOR Elda Araya & Carolina Cano Saint Kitts and Nevis 500 10.86472 Rio Blanco Abajo 1 -85.41531 18 469 50 23 27 GUANACASTE paratype 3428543394 2004-09-12 2017-10-02 2004-09-12 SECTOR Gloria Sihezar Saint Kitts and Nevis 500 10.86472 Rio Blanco Abajo 1 -85.41531 18 469 50 23 27 GUANACASTE paratype 3428568307 2010-11-11 2016-07-11 2010-11-11 SECTOR Lucia Rios Saint Kitts and Nevis 620 11.01373 Monte Cristo 1 -85.42531 Bosque Aguirre 18 469 17 3 14 GUANACASTE paratype 3428543320 2003-06-12 2015-01-25 2003-06-12 SECTOR Lucia Rios & Manuel Rios & Calixto Moraga Saint Kitts and Nevis Sendero Cuestona 640 10.98171 Sendero Laguna 1 -85.42785 Sendero Evangelista 18 469 81 32 49 GUANACASTE paratype 3428543326 2003-06-12 2015-01-25 2003-06-12 SECTOR Manuel Rios Freddy Quesada Saint Kitts and Nevis Sendero Cuestona 640 10.98171 Sendero Laguna 1 -85.42785 Sendero Evangelista 18 469 81 32 49 GUANACASTE paratype 3428543364 2003-06-12 2015-01-25 2003-06-12 SECTOR Lucia Rios & Manuel Rios & Calixto Moraga & Freddy Quesada & Petrona Rios Saint Kitts and Nevis Sendero Cuestona 640 19 470 10.98171 Sendero Laguna 1 -85.42785 Sendero Evangelista 18 469 81 32 49 GUANACASTE paratype 3428543377 2003-06-12 2015-01-25 2003-06-12 SECTOR Lucia Rios & Manuel Rios Saint Kitts and Nevis Sendero Cuestona 640 10.98171 Sendero Laguna 1 -85.42785 Sendero Evangelista 19 470 81 32 49 GUANACASTE paratype 3428543373 2003-06-12 2015-01-25 2003-06-12 SECTOR Manuel Rios & Freddy Quesada & Petrona Rios Saint Kitts and Nevis Sendero Cuestona 640 10.98171 Sendero Laguna 1 -85.42785 Sendero Evangelista 19 470 81 32 49 GUANACASTE paratype 3428543370 2011-09-27 2017-09-02 2011-09-27 USNM Freddy Quesada Saint Kitts and Nevis Sendero Nacho 710 10.9835 Sendero Tierra Blanca 5 -85.42117 Sendero Naciente 19 470 15 10 5 GUANACASTE paratype 3428543343 2011-09-27 2017-09-02 2011-09-27 USNM Manuel Rios Saint Kitts and Nevis Sendero Nacho 710 10.9835 Sendero Tierra Blanca 5 -85.42117 Sendero Naciente 19 470 15 10 5 GUANACASTE paratype 3428543341 2011-09-27 2017-09-02 2011-09-27 USNM Freddy Quesada & Manuel Rios & Calixto Moraga Saint Kitts and Nevis Sendero Nacho 710 10.9835 Sendero Tierra Blanca 5 -85.42117 Sendero Naciente 19 470 15 10 5 GUANACASTE paratype 3428543302 2011-09-27 2017-09-02 2011-09-27 USNM Manuel Rios Saint Kitts and Nevis Sendero Nacho 710 10.9835 Sendero Tierra Blanca 5 -85.42117 Sendero Naciente 19 470 15 10 5 GUANACASTE paratype 3428543354 2011-09-27 2017-09-02 2011-09-27 USNM Freddy Quesada Saint Kitts and Nevis Sendero Nacho 710 10.9835 Sendero Tierra Blanca 5 -85.42117 Sendero Naciente 19 470 15 10 5 GUANACASTE paratype 3428543369 2011-09-27 2017-09-02 2011-09-27 USNM Freddy Quesada Saint Kitts and Nevis Sendero Nacho 710 10.9835 Sendero Tierra Blanca 5 -85.42117 Sendero Naciente 19 470 15 10 5 GUANACASTE paratype