A revision of Scipopus Enderlein including the subgenera Scipopus s. str., Phaeopterina Frey and Parascipopus subgen. nov. (Diptera, Micropezidae, Taeniapterinae) Lindsay, Kate Marshall, Stephen A. European Journal of Taxonomy 2023 2023-11-08 904 1 189 C2NMJ Lindsay & Marshall, 2023 Lindsay & Marshall 2023 [462,902,683,710] Insecta Micropezidae Scipopus Animalia Diptera 88 89 Arthropoda species fenestratus subgen. et sp. nov. Parascipopus   urn:lsid:zoobank.org:act: 269ABDDC-F6F2-4B07-A791-227B5432BC5B  Figs 36–37    Differential diagnosis   Scipopus( Parascipopus) fenestratusresembles  S.( Pa.) nigriscapus sp. nov.in having a mostly orange body, but it differs by the three well-defined subapical hyaline spots on the wing, the absence of an apical scutellar seta, and the black distal and basal bands on the femora.    Etymology The species name, from the Latin noun for ‘window’, refers to the subapical well-defined, window-like hyaline spots on the wing.    Type material examined   Holotype   COSTA RICA• 1 ♀; Puntarenas, Monteverde Bio. Stn, along lower trail; 10°18′7″ N, 84°48′10″ W;  1500 ma.s.l.;  14 Aug. 2010; S.A. Marshallleg.; MNCR.   Paratypes   COSTA RICA• 2 ♀♀; same collection data as for holotype; MNCR• 1?; same collection data as for preceding; MYCRO015-15/UGIC114-15 sequenced for CO1; MNCR•  1 ♀; Puntarenas, Coto Brus, Z.P. Las Tablas, Est. Biol. Las Alturas; 82°50′04″ W, 08°57′07″ N;  1500–1600 ma.s.l.;  18–24 Jun. 2013; ProjectoZADBI; Malaise Trap; ZADBI -889; MNCR• 1 ♂; San José, Moravia, Zurquí de Moravia, Northpasture; 84°00′57″ W, 10°02′58″ W;  1600 ma.s.l.;  22–29 Mar. 2013; ProyectoZADBI; Malaise trap#3; ZADBI-662; MNCR•  1 ♀; same collection data as for preceding;  12–19 Apr. 2013; ZADBI-712; MYCRO045-15/UGIC144-15 sequenced for CO1; MNCR•  1 ♀; same collection data as for preceding; Creek2 north;  24–31 May 2013; Malaise trap#2; ZADBI-786; MNCR(photographed, Fig. 36C)•  1 ♀; same collection data as for preceding;  10–17 May 2013; ZADBI-743; MNCR•  1 ♀; Prov. Cartago, Paraíso, P.N. Tapanti-Macizo de la Muerte;  1420 ma.s.l.;  19 May 2000; C. Hansonand D. Rubileg.; Malaise; L_N_188500_561800 #56722; MNCR• 1 ♂; San José, est. Zurqui  500 mantes de Tunel;  1600 ma.s.l.;  Mar. 1991; G. Maassleg.; L-N-226800, 535200; MNCR.   Other material examined   COSTA RICA• 1 ♀; Prov. San José, Moravia, Zurquí de Moravia, Towerpath; 10°02′58″ N, 84°00′57″ W;  1600 ma.s.l.;  12–19 Apr. 2013; ProjectoZADBI; Malaise trap#1; ZADBI-710; MNCR•  1 ♀; same collection data as for preceding;  24–31 May 2013; ZADBI-785; MNCR•  1 ♀; same collection data as for preceding; Northpasture; Malaise trap#3; ZADBI-787; MNCR•  1 ♀; same collection data as for preceding;  16Apr. 2013; hand collected; ZADBI-663; MNCR•  1 ♀; same collection data as for preceding; Creek2 north;  12–19 Apr. 2013; Malaise trap#2; ZADBI-711; MNCR•  1 ♀, 1?; same collection data as for preceding;  5–9 Aug. 2013; S.A. Marshallleg.; hand collected at light; ZADBI-1193; MNCR•  2 ♀♀; same collection data as for preceding;  5–6 Aug. 2013; mixed methods; ZADBI-1041; MNCR•  1 ♀; same collection data as for preceding;  20 Oct.–1 Nov. 2012; ProjectoZADBI; Malaise trap#1; ZADBI-190; MNCR•  1 ♀; same collection data as for preceding; Creek2 north;  24–31 May 2013; Malaise trap#2; ZADBI-786; MNCR•  1 ♀; Prov. Cartago, Paraiso, P.N. Tapanti-Macizo de la Muerte;  1500 ma.s.l.  19 May 2000; C. Hansonand D. Rubileg.; Malaise; TorreICE #28; L_N_189400_561700; MNCR(dissected and photographed, Fig. 36A).    Description  LENGTH. 13–15 mm.  HEAD. Antenna orange. Palpus orange, pale orange microtrichose and sparsely black setulose on entire surface, broad (length 3.1× height). Clypeus orange, width ~2.4× height, silvery microtrichose on entire surface. Frontal vitta entirely dull, orange, microtrichose, tapered posteriorly but posterior apex expanded. Orbital plate dull, orange, microtrichose, clearly delineated. Epicephalon orange, shiny, white-microtrichose, wide (width>⅔ of frontal vitta width at inner verticals), clearly delineated from upper frontal vitta. Paracephalon orange or light brown. One lower fronto-orbital seta, upper fronto-orbital seta absent or present, outer vertical seta absent, all other head chaetotaxy well-developed.  THORAX. Scutum orange, microtrichose. Female cervical sclerite convex. Postpronotal lobe orange, orange setulose anteriorly and on outer margin. Notopleuron orange. Pleuron orange, uniformly white microtrichose. Tibiae dark brown or black, fore femur orange on basal ¾, dark brown or black apically, mid and hind femora pale brown or orange basally and preapically, darker brown or black medially, black apically. First fore tarsomere yellow, hind tarsomere 1 dark brown or black with ventral golden fringe. Wing brown infuscate on r 1, distal ½ and apex; with three hyaline window-like spots in m 1, r 2+3, r  3+4( Fig. 37A).  ABDOMEN(J+ ♀). Tergites orange or brown, black setulose. T1 with long, fine, yellow or orange setae. Pleuron yellow or pale orange, microtrichose, dark orange or brown on P1, dorsal half of P2–6 dark orange ( Fig. 37B).  FEMALE ABDOMEN. T1+2 ~2.2× length of T3. Oviscape orange and/or brown, white microtrichose on anterior ½, ~4.0–5.0 × length of T6. Combined spermathecal duct length short; common duct ½ of entire duct length. Paired spermathecal duct very short (1.0–2.0 × length of paired spermathecae), very swollen medially, less swollen apically. Paired spermathecal stems inornate, bulbous and wrinkled, longer than spermathecae. Paired spermathecae spherical, with apical indentation (collapsed in Fig. 36A). Single spermathecal duct arising from base of paired duct, narrow, ½ the diameter and ≈ length to paired duct. Single spermatheca swollen distally, inornate.  MALE ABDOMEN. T1+2 ~3.0× length of T3. T6 short, genital fork 2.3 × as long as T6, arms converging, inner basal process angled inwards, ¹/5the length of arm. Epandrium elongate, length ~2.0× height, short setose on posteroventral margin. Basiphallus small, crescent-shaped. Basal distiphallus long (~½ epandrial length), ending in small, short (length ≈ height) single-chambered phallic bulb, with narrow anterior projections. Distal distiphallus narrow, long (≈ length to epandrium), with apex slightly expanded and tube-like. Phallapodeme broad and expanded apically. Hypandrium expanded anteriorly.    Distribution  Costa Rica. 4465943756 2010-08-14 MNCR S. A. Marshall Costa Rica 1500 10.301945 Monteverde Bio. Stn, along lower trail 21 -84.80278 88 89 1 1 Puntarenas holotype 4465943571 [189,1034,1380,1406] 2010-08-14 MNCR S. A. Marshall Costa Rica 1500 10.301945 Monteverde Bio. Stn, along lower trail 21 -84.80278 88 89 2 2 Puntarenas paratype 4465943537 2010-08-14 MNCR S. A. Marshall Costa Rica 1500 10.301945 Monteverde Bio. Stn, along lower trail 21 -84.80278 88 89 1 1 Puntarenas paratype 4465943541 2013-06-18 2013-06-24 2013-06-18 Malaise trap MNCR Costa Rica 1550 8.951944 Z.P. Las Tablas, Est. Biol. Las Alturas 21 -82.83445 Coto Brus 88 89 1 1 Puntarenas paratype 4465943357 2013-03-22 2013-03-29 2013-03-22 Malaise trap MNCR Costa Rica 1600 Zurquí de Moravia, North pasture -10.049445 Moravia 88 89 1 1 San Jose paratype 4465943738 2013-04-12 2013-04-19 2013-04-12 Malaise trap MNCR Costa Rica Moravia 1600 Zurquí de Moravia, North pasture -10.049445 Moravia 88 89 1 1 San Jose paratype 4465943619 2013-05-24 2013-05-31 2013-05-24 Malaise trap MNCR Costa Rica 1600 Zurquí de Moravia, North pasture, Creek 2 north -10.049445 Moravia 88 89 1 1 1 San Jose paratype 4465943960 [332,1327,1668,1695] 2013-05-10 2013-05-17 2013-05-10 Malaise trap MNCR Costa Rica 1600 Creek -10.049445 Moravia 88 89 1 1 1 San Jose paratype 4465943726 2000-05-19 Malaise MNCR C. Hanson & D. Rubi Costa Rica 1420 P.N. Tapanti-Macizo de la Muerte Paraíso 88 89 1 1 Cartago paratype 4465943519 1991-03-31 MNCR G. Maass Costa Rica 500 est. Zurqui 500 m antes de Tunel Zurqui 88 89 1 1 San Jose paratype 4465943709 2013-04-12 2013-04-19 2013-04-12 MNCR Costa Rica 1600 10.049445 Zurqui de Moravia, Tower path 21 -84.01583 Moravia 88 89 1 1 San Jose 4465943554 2013-05-24 2013-05-31 2013-05-24 MNCR Costa Rica 1600 10.049445 Zurqui de Moravia, Tower path 21 -84.01583 Moravia 88 89 1 1 San Jose 4465943424 2013-05-24 2013-05-31 2013-05-24 Malaise trap MNCR Costa Rica 1600 10.049445 Zurqui de Moravia, North pasture 21 -84.01583 Moravia 88 89 1 1 San Jose 4465943373 2013-04-16 2013-05-31 2013-05-24 Malaise trap MNCR Costa Rica 1600 90 91 10.049445 Zurqui de Moravia, North pasture 21 -84.01583 Moravia 88 89 1 1 San Jose 4465943530 2013-04-12 2013-04-19 2013-04-12 Malaise trap MNCR Costa Rica 1600 10.049445 Zurqui de Moravia, Creek two north 21 -84.01583 Moravia 90 91 1 1 San Jose 4465943591 2013-08-05 2013-08-09 2013-08-05 hand collected at light MNCR S. A. Marshall Costa Rica 1600 10.049445 Zurqui de Moravia, Creek two north 21 -84.01583 Moravia 90 91 2 1 San Jose 4465943974 [191,1379,370,397] 2013-08-05 2013-08-06 2013-08-05 mixed methods MNCR S. A. Marshall Costa Rica 1600 10.049445 Zurqui de Moravia, Creek two north 21 -84.01583 Moravia 90 91 2 2 San Jose 4465943701 2012-10-20 2012-11-01 2012-10-20 Malaise trap MNCR S. A. Marshall Costa Rica 1600 10.049445 Zurqui de Moravia, Creek two north 21 -84.01583 Moravia 90 91 1 1 San Jose 4465943858 2013-05-24 2013-05-31 2013-05-24 Malaise trap MNCR S. A. Marshall Costa Rica 1600 10.049445 Creek 21 -84.01583 Moravia 90 91 1 1 San Jose 4465943719 2000-05-19 Malaise MNCR C. Hanson & D. Rubi Costa Rica 1500 P.N. Tapanti-Macizo de la Muerte Paraiso 90 91 1 1 Cartago