A revision of Scipopus Enderlein including the subgenera Scipopus s. str., Phaeopterina Frey and Parascipopus subgen. nov. (Diptera, Micropezidae, Taeniapterinae) Lindsay, Kate Marshall, Stephen A. European Journal of Taxonomy 2023 2023-11-08 904 1 189 C2NMG Lindsay & Marshall, 2023 Lindsay & Marshall 2023 [459,905,850,877] Insecta Micropezidae Scipopus Animalia Diptera 97 98 Arthropoda species monteverde subgen. et sp. nov. Parascipopus   urn:lsid:zoobank.org:act: 22CDF6EE-0943-4699-993A-EC3FE6D0E7C8  Fig. 41    Differential diagnosis   Scipopus( Parascipopus) monteverde sp. nov.resembles  S.( Pa.) tico sp. nov.in having an orange frontal vitta and a primarily dark brown or black body with a blue sheen, but it differs in having evenly dark brown or black femora, a darkly infuscate wing, and a dark brown or black first fore tarsomere.    Etymology The name is a noun in apposition inspired by the typelocality.    Type material examined   Holotype   COSTA RICA• 1 ♀; Monteverde, sweep nr Biol Stn;  25 May 1998; S.A. Marshallleg.; pseudoscorpion associated with specimen; MNCR(photographed, Fig. 41F).   Paratypes   COSTA RICA• 1 ♂; Prov. Puntarenas, Coto Brus, Z.P. Las Tablas, Est. Biol. Las Alturas; 82°50′04″ W, 08°57′07″ N;  1500–1600 ma.s.l.;  2–9 Apr. 2013; ProjectoZADBI; Malaise Trap; ZADBI -683; MYCRO170-15/UGIC169-15 sequenced for CO1; MNCR•  1 ♀; same collection data as for preceding; MNCR•  1 ♀; same collection data as for preceding;  15–22 Apr. 2013; ZADBI685; MNCR•  1 ♀; same collection data as for preceding;  13–20 May 2013; ZADBI-884; MNCR•  1 ♀; same collection data as for preceding;  22–29 Apr. 2013; ZADBI-881; MYCRO166-15 UGIC265-15 sequenced for CO1; MNCR(photographed, Fig. 41A)• 1 ♂; same collection data as for preceding;  5–11 Mar. 2013; ZADBI-679• 1 ♂; Prov. Alajuela, Upala, Albergue Heliconias, S. Laguna Danto;  1100 ma.s.l.;  24–26 Jan. 2001; J.D. Gutiérrezleg.; manual; L_N_423760_298100; MNCR• 1 ♂; Prov. Puntarenas, Buenos Aires, Estación Altamira, Sendero Los Gigantes;  1460 ma.s.l.;  2–18 Feb. 2001; O. Alemánleg.; manual; L_S_572200_331700; MNCR• 1 ♂; Prov. Cartago, Turriabla, Santa Teresita, Monumento Nacional Guayabo;  1100 ma.s.l.;  Oct. 1994; G. Fonsecaleg.; Malaise; L_N_570000_217400; MNCR•  1 ♀; Puntarenas, Monteverde Biol. Res.;  1500 ma.s.l.;  12 Jun. 2000; M. Buckleg.; cloud forest; debu00118150; DEBU•  2 ♀♀; same locality as for preceding; lower trail;  11 Jun. 2000; S.A. Marshallleg.; debu00118248; DEBU•  1 ♀; same locality as for preceding;  1500 ma.s.l.;  13–14 Jun. 2000; D. Bickelleg.; sticky trapon tree trunk; debu00138177; MNCR(dissected and photographed, 41C)•  1 ♀; same locality as for preceding; along lower trail; 10°18′7″ N, 84°48′10″ W; ~  1500 ma.s.l.;  14 Aug. 2012; S.A. Marshallleg.; debu0033024; MNCR(photographed, Fig. 41D)• 1 ♂; San José, SanCarlos, Cerro Cura,  14 kmSSW of San Marcos de Tarrazú;  1800 ma.s.l.; 9°36′31″ N, 84°7′9″ W;  15 Feb. 2003; S.A. Marshallleg.; debu00205682; DEBU(dissected and photographed, Fig. 41B, E).    Description  LENGTH. 10–13 mm.  HEAD. Antenna orange. Palpus dark brown, apically light brown and pointed, pale microtrichose and setulose, narrow (length 4.1 × height). Clypeus dark brown or black, width ~2.1× height, silvery microtrichose on entire surface. Frontal vitta dull, orange, microtrichose, posterior apex strongly tapered to a point ( Fig. 41D). Orbital plate orange, shiny, bare, clearly delineated. Epicephalon black or dark brown, shiny, bare, wide (width>⅔ of upper frontal vitta width at inner verticals), clearly delineated from upper frontal vitta. Paracephalon dark brown or black. Upper fronto-orbital seta and one of the usual two lower fronto-orbital setae absent, all other head chaetotaxy well-developed.  THORAX. Scutum black, brown microtrichose, with a wide, shiny, blue indistinct sheen medially. Female cervical sclerite slightly convex. Postpronotal lobe black, setulose anteriorly. Notopleuron black with blue sheen at some angles. Pleuron black with blue sheen, entirely covered in silvery microtrichosity, denser on posterior katepisternum, forming silvery transverse band only seen at some angles ( Fig. 41F). Legs black, mid and hind femora sometimes slightly paler basally; fore and hind first tarsomere black with ventral golden fringe. Wing slightly brown infuscate. Apical scutellar seta absent.  ABDOMEN(J+ ♀). T1 with fine, long, white setae.  FEMALE ABDOMEN. Pleuron mostly dark brown or black, ventral half of P1–3 grey ( Fig. 41G). T1+2 ~2.2 × length of T3. Oviscape black, setulose, white microtrichose on anteroventral and lateral ⅓, dorsomedian ¹/6; ~3.0× length of T6. Common duct very short, ⅛ of entire duct length. Paired spermathecal duct long (7.0 × length of paired spermathecae), mostly parallel-sided, wider basally, swollen apically. Paired spermathecal stems separated into very short, narrow basal section and long, swollen, distal section; sparse tubercles at base of paired spermathecae. Paired spermathecae cup-like, with apical indentation. Single spermathecal duct arising from basal ¹/6of common duct, narrow, ½ diameter of base of paired duct, <½ the length of paired duct, slightly swollen apically. Single spermatheca teardrop-shaped, elongate.  MALE ABDOMEN. Pleuron pale grey or off-white, dark brown or black on P1, dorsal ¾ of pleural sac and dorsal half of P2–6. T1+2 ~2.6× length of T3. Genital fork 2.0× as long as T6, arms converging; inner basal process angled inwards, ¹/6the length of arm. Epandrium elongate (length ~2.0× height), long setose on posteroventral margin. Basiphallus small, projecting posteriorly, crescent-shaped. Basal distiphallus long (> ½ epandrial length), partially ending in large, long (length 2.0× height), approximately ovoid phallic bulb. Distal distiphallus wide, long (≈ length to epandrium, diameter slightly greater than basal distiphallus). Phallapodeme apically expanded and broad (not visible in Fig. 41E). Anterior hypandrium expanded.    Remarks A female  S.( Pa.) monteverde sp. nov.was photographed ovipositing in mossy substrate in Monteverde, Costa Rica( Fig. 41G).    Distribution  Costa Rica. 4465943423 1998-05-25 MNCR S. A. Marshall Costa Rica sweep nr Biol Stn Monteverde 97 98 1 1 holotype 4465943368 2013-04-02 2013-04-09 2013-04-02 Malaise trap MNCR Costa Rica 1550 8.951944 Z. P. Las Tablas, Est. Biol. Las Alturas 21 -82.83445 Coto Brus 97 98 1 1 Puntarenas paratype 4465943919 2013-04-02 2013-04-09 2013-04-02 Malaise trap MNCR Costa Rica 1550 8.951944 Z. P. Las Tablas, Est. Biol. Las Alturas 21 -82.83445 Coto Brus 97 98 1 1 Puntarenas paratype 4465943795 [308,1310,1608,1635] 2013-04-15 2013-04-22 2013-04-15 MNCR Costa Rica 1550 8.951944 Est. Biol. Las Alturas 21 -82.83445 Prov. 97 98 1 1 Puntarenas paratype 4465943602 2013-05-13 2013-05-20 2013-05-13 Malaise trap MNCR Costa Rica 1550 8.951944 Z. P. Las Tablas, Est. Biol. Las Alturas 21 -82.83445 Coto Brus 97 98 1 1 Puntarenas paratype 4465943669 2013-04-22 2013-04-29 2013-04-22 MNCR Costa Rica 1550 8.951944 Est. Biol. Las Alturas 21 -82.83445 Prov. 97 98 1 1 Puntarenas paratype 4465943822 2013-03-05 2013-03-11 2013-03-05 Malaise trap MNCR Costa Rica 1550 8.951944 Z. P. Las Tablas, Est. Biol. Las Alturas 21 -82.83445 Coto Brus 97 98 1 1 Puntarenas paratype 4465943675 2001-01-24 2001-01-26 2001-01-24 MNCR J. D. Gutierrez Costa Rica 1100 Albergue Heliconias, S. Laguna Danto Upala 97 98 1 1 Alajuela paratype 4465943936 2001-02-02 2001-02-18 2001-02-02 MNCR O. Aleman Costa Rica 1460 Estación Altamira, Sendero Los Gigantes Buenos Aires 97 98 1 1 Puntarenas paratype 4465943708 1994-10 Malaise MNCR G. Fonseca Costa Rica 1100 Santa Teresita, Monumento Nacional Guayabo Turriabla 97 98 1 1 Cartago paratype 4465943740 2000-06-12 DEBU M. Buck Costa Rica 1500 Monteverde Biol. Res. 97 98 1 1 Puntarenas paratype 4465943559 2000-06-11 DEBU S. A. Marshall Costa Rica 1500 Monteverde Biol. Res. lower trail 97 98 2 2 Puntarenas paratype 4465943607 2000-06-13 2000-06-14 2000-06-13 MNCR D. Bickel Costa Rica 1500 98 99 Monteverde Biol. Res. lower trail 97 98 1 1 Puntarenas paratype 4465943842 2012-08-14 2000-06-14 2000-06-13 MNCR S. A. Marshall Costa Rica 1500 10.301945 Monteverde Biol. Res. along lower trail 21 -84.80278 98 99 1 1 Puntarenas paratype 4465943759 2003-02-15 DEBU S. A. Marshall Costa Rica 1800 9.608611 Cerro Cura, 14 km SSW of San Marcos de Tarrazú 21 -84.11917 San Carlos 98 99 1 1 San Jose paratype