Actinopus palmar Ríos-Tamayo & Goloboff, 2018

Miglio, Laura Tavares, Pérez-Miles, Fernando & Bonaldo, Alexandre B., 2020, Taxonomic Revision of the Spider Genus Actinopus Perty, 1833 (Araneae, Mygalomorphae, Actinopodidae), Megataxa 2 (1), pp. 1-256 : 207-210

Actinopus palmar Ríos-Tamayo & Goloboff, 2018

Figs 187–189 , Map 11

Actinopus palmar Ríos-Tamayo & Goloboff, 2018: 81 , fig. 26 A–G, 45 (holotype ♂, [31º 51’ S 58º 19’ W], Parque Nacional “El Palmar”, Entre Ríos, 12–16.ii.1980, P. Grosso and P. Goloboff leg., MACN-Ar 198206, examined); World Spider Catalog, 2020.

Diagnosis. Males of Actinopus palmar differ from those of all other species of the group cucutaensis by the narrow ATP base, copulatory bulb with three prolateral keels in prolaterodorsal view and ATP with a big serrated area ( Fig. 189 A–C). Additionally, they differ from those of all other species of Actinopus , except A. crassipes , A. dubiomaculatus , A. pusillus , A. emas , A. bocaina , A. laventana and A. gerschiapelliarum , by the distal leg articles being paler than proximal leg articles and the palp.

MALE (MACN 19826): Total length 8.62; Carapace, long 4.3; wide 3.6. Carapace anterior part tapering. Eyes. Anterior row slightly procurved, posterior row recurved ( Fig. 187 B ). One bristle between AME-clypeus and also ALE–PLE. Sternum with eight sigilla near center and two distal sigilla very fused to each other ( Fig. 187 C ). Rastellum protuberant, inverted V-shaped, hirsute, with many spines on apex ( Fig. 189 D–E). Chelicerae with denticles along Prolateral row ( Fig. 187 D ). Patella and tibia III with distal crown of well-developed thorns ( Fig. 188 A–B).Patellae III and IV with spines on prolaterodorsal face ( Fig. 188 A, C ). Patella IV without spines dorsally ( Fig. 188 D ). Ventral pseudoscopulae occupying 30% of tarsi I, 50% of II and 100% of III and IV. Pseudoscopula of tarsi I and II with setae spaced, forming diffuse group of bristles; On tarsi III and IV, compact. Carapace and Chelicerae reddish brown, sternum yellowish brown, coxae, trochantera, femora and patellae reddish brown, tibiae, metatarsi and tarsi yellowish brown, abdomen pale gray ( Fig. 187 A ). Eyes: Diameters: PME 0.15 PLE 0.18 ALE 0.28, AME 0.21; MOQ: Length 0.68, front width 1.78, back width 1.5; Interdistances: PME–PME 0.90, PLE–PME 0.12, AME–AME 0.09, ALE–AME 0.65, ALE–PLE 0.34, AME–PME 0.25. Ocular area: OAL 2.53, OAW 2.62, and IF 1.4. Body: Clypeus: 0.15; Fovea: 1.3; Labium: long 0.85; wide 0.75; Chelicerae: long 2.43; wide 1.18; Sternum: long 2.8; wide 2.1. Abdomen: long 4.56; wide 3.5. Leg measurements: I: Fe 3.62/ Pa 1.62/ Ti 2.12/ Me 3.25/ Ta 1.75/ total 12.37. II: 4/ 1.75/ 2.5/ 2.87/ 1.87/ 13. III: 3/ 2.37/ 1.75/ 3/ 2.125/ 12.25. IV: 4.12/ 1.37/ 3.25/ 3.12/ 2.37/ 14.25. Formula 4123. Spination: I—Fe v0, d0, p0, r0; Pa v0-0-1, d0, p0, r0; ti v5-6-8, d0, p0, r0; Me v5-14-11, d0, p1-0-2, r0; ta v6-10-9, d0, p0-0- 2, r1-3-4. II—Fe v0, d0, p0, r0; Pa v0-0-2, d0, p0, r0; ti v4-4-6, d0, p0-0-1, r4-8-6; Me v5-5-8, d0, p0-0-2, r2-2-4; ta v7-11-13, d0, p0-0-1, r3-3-7. III—Fe v0, d1-4-3, p0, r0; Pa v0-0-2, d5-3-14, p2-0-3, r0-0-2; ti v0-0-2, d1-0-9, p0- 0-3, r0-0-2; Me v0-0-3, d1-3-0, p0-2-1, r0-1-2; ta v0, d0, p1-2-7, r0-2-5; IV—Fe v0, d0, p0, r0; Pa v0, d16-16-8, p0-1-1, r0; ti v0-0-2, d0, p0-0-1, r0; Me v0-0-2, d0, p0-2- 3, r0; ta v0, d0, p2-6-7, r0-1-2. Palp: PA well-developed embolus with three keels (Pa, PI and PS). ATP almost 1/2 size of embolus. BTA developed; serrated area large, from beside embolus base to middle of embolus; embolus inserted at 80°; embolar base thin ( Fig. 189 A–C).

VARIATION

MALES (n=2): Eyes: Diameters: PME 0.12–0.15, PLE 0.15–0.18, ALE 0.28–0.31, AME 0.12–0.21; MOQ: Length 0.62–0.68, front width 1.68–1.78, back width 1.5– 1.56; I nterdistances: PME–PME 0.84–0.90, PLE–PME 0.09–0.12, AME–AME 0.09–0.15, ALE–AME 0.59–0.65, ALE–PLE 0.28–0.34, AME–PME 0.25–0.28. Ocular area : OAL 2.5–2.53, OAW 2.12–2.62, and IF 1.12–1.4. Body: Total length: 6.75–8.62; Carapace: long 3.62–4.3; wide 3.25–3.6; Clypeus: 0.12–0.15; Fovea: 1.30–1.40; Labium: long 0.84–0.85; wide 0.75–0.76; Chelicerae: long 1.56–2.43; wide 1.03–1.18; Sternum: long 2.24–2.8; wide 2–2.1. Abdomen: long 3.62–4.56; wide 2.75–3.5. Leg measurements: I: Fe 3.62–4/ Pa 1.37–1.62/ Ti 2.12– 2.37/ Me 2.87–3.25/ Ta 1.75–1.87/ total 12.37–12.5. II: 3.75–4/ 1.12–1.75/ 2.12–2.5/ 2.87/ 1.75–1.87/ 11.62–13. III: 2.75–3/ 1.5–2.37/ 1.25–1.75/ 2.5–3/ 2–2.12/ 10–12.25. IV: 3.62–4.12/ 1.37–1.62/ 2.75–3.25/ 3–3.12/ 2.25–2.37/ 13.25–14.25. Formula 4123 .

Material examined. ARGENTINA: Entre Ríos, Parque Nacional “El Palmar”, [58° 19’ 21” S 31° 51’ 11” W], 1♂, 12–16.ii.1980, Grosson & P. A. Goloboff leg. ( MACN 19826 ) ; iii.1995, C. J. Grismado leg. ( MACN 19839 ) .

Distribution. ARGENTINA. Entre Ríos ( Map 11 ).

Figures

MAP 11. Distribution map of Actinopus robustus O. Pickard-Cambridge, 1892, Actinopus guajara sp. nov., Actinopus palmar Ríos-Tamayo & Goloboff, 2018, Actinopus lomalinda sp. nov., Actinopus concinnus sp. nov. and Actinopus cucutaensis (Mello- Leit„o, 1941).

FIGURE 187. Actinopus palmar Ríos-Tamayo & Goloboff, 2018, male:A–D MACN-Ar 19826:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral.

FIGURE 188. Actinopus palmar Ríos-Tamayo & Goloboff, 2018, male: 815–818 MACN-Ar 19826:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal.

FIGURE 189. Actinopus palmar Ríos-Tamayo & Goloboff, 2018, male: A–E MACN 19826: A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; BTA-Basal Tegular Apophysis; PAc- Prolateral Accessory keel; PI-Prolateral Inferior keel; PS-Prolateral Superior keel; Arrows: Serrated area. E. Rastellum, dorsal; F. Rastellum, ventral. Scales lines: A–C: 1 mm; D–E: 0.5 mm.