EPITONIIDAE Berry, 1910

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 610-612

Family EPITONIIDAE Berry, 1910 [1812]

Acyonaea – Deshayes, 1830a-EncyMeth: 6, ex Leach ms. Incorrect subsequent spelling of Aciona Leach, 1815 , an objective synonym of Epitonium Röding, 1798 .

aemula , Scalaria – Deshayes, 1861-DescrCoqFoss: 348, 14, pl. 14, figs 7-9. — Type localities:Châlons-sur-Vesle & Jonchery-sur-Vesle.— Type age: Paleocene (Thanetian). — Current status: Opalia (Crassiscala) aemula (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) .

affinis , Scalaria –Deshayes, 1861-DescrCoqFoss: 339-340, 12, pl. 12, figs 4-5. — Type localities: Le Guépelle & Chéry-Chertreuve. — Type age: Eocene (Bartonian). — Current status: Amaea (Acrilla) affinis (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) .

aizyensis , Scalaria –Deshayes, 1861-DescrCoqFoss: 332-333, 12, as Scalaria “ multicincta Watelet ”, pl. 12, fig. 22, 13, 13, pl. 13, figs 14- 16 [no correction noted, so figure on pl. 12 uncertain]. — Type localities: Aizy-Jouy, Cuise-la-Motte & Laon. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32416, syntypes from Cuise-la-Motte. — Current status: Epitonium (Crisposcala) aizyensis (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) .

angusta , Scalaria – Deshayes, 1861-DescrCoqFoss: 342, 12, pl. 12, figs 6-7 [22 May]; 1861-DescrCoqFoss: 342 [20 November]. Not preoccupied by Scalaria angusta Dunker, 1861 [pre-August]. — Type localities: Thiverval-Grignon & Berchères-sur-Vesgre. — Type age: Eocene (Lutetian). Scala (Acrilla) perangusta de Boury, 1913 (de Boury 1913: 87, 96), unnecessary replacement name. Amaea (Acrilla) perangusta ( Boury, 1913) ( Le Renard & Pacaud 1995: 93) . — Current status: Acrilla angusta (Deshayes, 1861) (Pacaud herein).

auversiensis , Scalaria – Deshayes, 1861-DescrCoqFoss: 345, 25, pl. 25, fig. 1. — Type localities: Auvers-sur-Oise & Mary-sur-Marne. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32487, syntype from Auvers-sur-Oise. — Current status: Acirsa (Plesioacirsa) auversiensis (Deshayes, 1861) ( Le Renard & Pacaud 1995: 94) .

barrandei , Scalaria –Deshayes, 1861-DescrCoqFoss: 332, 12, pl. 12, figs 11-12. — Type locality: Cuise-la-Motte. — Type age: Eocene (Ypresian). — Current status: Epitonium (Crisposcala) barrandei (Deshayes, 1861) ( Le Renard & Pacaud 1995: 92) .

brevicula , Scalaria –Deshayes, 1861-DescrCoqFoss: 335, 11, pl. 11, figs 13-14. — Type locality: Le Fayel. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32406, 2 syntypes. — Current status: Cirsotrema (Circuloscala) breviculum (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) .

caelata , Melania – Deshayes, 1862-DescrCoqFoss: 452, 30, pl. 30, figs 1-3. — Type locality: Bazoches. — Type age: Eocene (Ypresian). Acirsa (Plesioacirsa) subtenuistriata [d’Orbigny, 1850] caelata (Deshayes, 1862) ( Le Renard & Pacaud 1995: 94) . — Current status: synonym of Acirsa (Plesioacirsa) subtenuistriata (d’Orbigny, 1850) (Pacaud 2007: 42) .

caillati , Scalaria – Deshayes, 1861-DescrCoqFoss: 351, 13, pl. 13, figs 18-20. —Type localities: Thiverval-Grignon, Parnes & Damery.— Type age: Eocene (Lutetian). — Type material: UCBL-EM 32424, syntype from Thiverval-Grignon. — Current status: synonym of Opalia (Crassiscala) francisci (Caillat, 1835) (Pacaud herein). contabulata , Scalaria – Deshayes, 1861-DescrCoqFoss: 334-335, 11, pl. 11, figs 11-12. — Type locality: Cuise-la-Motte. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32405, syntype. — Current status: Cirsotrema (Gyroscala) contabulatum (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) .

coronalis, Scalaria – Deshayes, 1861-DescrCoqFoss: 337-338, 11, as Scalaria “ coronata ”, pl. 11, figs 7-8; 1865-DescrCoqFoss: 666 [error corrected]. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32403, syntype. — Type species (OD) of Coroniscala de Boury, 1909. — Current status: Cirsotrema (Coroniscala) coronale (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93; Pacaud 2008: 58).

crassicostata , Scalaria – Deshayes, 1850-TraitElem: 42, pl. 70, figs 1-3, non G. B. Sowerby II, 1844. — Type locality: none given [ France]. — Type age: Miocene (Burdigalian). This species was a nomen nudum in Verneuil (1839: 76). Synonym of Scalaria (Cirsotrema) rustica Defrance, 1827 ( Boury 1913: 100). Cirsotrema crassicostatum (Deshayes, 1850) ( Cossmann & Peyrot 1922: 151- 152, pl. 4, figs 79-80, 97-98; Lozouet et al., 2001: 51-52, pl. 19, fig. 6). There are two other potential synonyms: Cirsotrema rustica (Defrance, 1827) [ Scalaria ]; Boury, 1913, considered it to be the same as Deshayes’ species, but this taxon was unfigured and the type was lost in the war in 1944 from the Defrance collection in Caen, and Cirsotrema thais Boury , in Cossmann, 1912, considered by Lozouet et al. (2001: 51-52) as a synonym of the Deshayes species. — Current status: Cirsotrema crassicostatum (Deshayes, 1850) ( Van der Voort 2024) .

elegantissima , Scalaria – Deshayes, 1861-DescrCoqFoss: 337, 12, pl. 12, figs 1-2. — Type localities: Le Vivray & Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Type species (OD) of Elegantiscala Boury, 1911. Family placement of Elegantiscala remains uncertain (L. G. Brown & Neville 2015: 23). — Current status: Cirsotrema (Elegantiscala) elegantissima (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93; Pacaud 2008: 58).

erasa , Scalaria – Deshayes, 1861-DescrCoqFoss: 350, 12, pl. 12, fig. 13, as S. “ erassa ”. Unnecessary replacement name for Melania tenuicosta Baudon, 1853 , non Scalaria tenuicostata Michaud, 1829 , which are not homonyms. — Type localities: Chaussy & Mouchy-le-Châtel. — Type age: Eocene (Lutetian). Acirsa (Acirsella) erasa (Deshayes, 1861) ( Le Renard & Pacaud 1995: 94; Pacaud 2008: 58). — Current status: Acirsa tenuicosta ( Baudon, 1853) (Pacaud herein).

gouldi , Scalaria – Deshayes, 1861-DescrCoqFoss: 346, 11, pl. 11, figs 15-16. — Type localities: Parnes & Saint-Félix. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32407, syntype from Parnes. — Type species (OD) of Pliciscala Boury, 1887. Opalia (Pliciscala) gouldi (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) . Opalia (Dentiscala) gouldi (Deshayes, 1861) ( Pacaud 2008: 58) . — Current status: Opalia (Pliciscala) gouldi (Deshayes, 1861) (Pacaud herein).

inermis , Scalaria (Pyrgiscus) – Deshayes, 1861-DescrCoqFoss: 354, 16, pl. 16, figs 26-28. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32447, 2 syntypes. — Type species (OD) of Acirsella Boury, 1886. — Current status:synonym of Acirsa canicularis (Lamarck, 1804) (Pacaud herein).

involuta, Scalaria – Deshayes, 1861-DescrCoqFoss: 333-334, 12, pl. 12, fig. 21. — Type localities: Cuise-la-Motte, Mercin-et-Vaux & Laon. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32415, syntype from Cuise-la-Motte. — Current status: synonym of Epitonium (Crisposcala) aizyense (Deshayes, 1861) (Pacaud herein).

lamarckii , Scalaria – Deshayes, 1861-DescrCoqFoss: 347, 11, pl. 11, figs 33-34. — Type locality: Hérouval. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32412, syntypes. Opalia (Pliciscala) lamarckii (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) . — Current status: synonym of Opalia (Pliciscala) monilifera (Melleville, 1843) (Pacaud herein).

lamberti , Scalaria – Deshayes, 1861-DescrCoqFoss: 349, 11, pl. 11, figs 27-28. — Type locality: Mercin-et-Vaux. — Type age: Eocene (Ypresian). — Type species (OD) of Torquatiscala Boury, 1912 , which has recently been regarded as a synonym of Cylindriscala Boury, 1909 . Amaea (Acrilla) lamberti (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) . — Current status: Cylindriscala l amberti (Deshayes, 1861) (Pacaud herein).

marginalis , Scalaria – Deshayes, 1861-DescrCoqFoss: 347, 11, pl. 11, figs 29-30. — Type localities: Cuise-la-Motte & Laon. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32409, syntype from Laon. — Current status: Opalia (Pliciscala) lamarckii [Deshayes, 1861] marginalis (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) .

michelini , Scalaria – Deshayes, 1861-DescrCoqFoss: 341, 23, pl. 23, figs 10-12. — Type locality: Le Fayel. — Type age: Eocene (Bartonian). — Current status: Tenuiscala (Tenuiscala) michelini (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) .

obsoleta , Scalaria – Deshayes, 1861-DescrCoqFoss: 348, 12, pl. 12, fig. 10. — Type locality: Aizy-Jouy. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32413, syntype. Synonym of Opalia (Pliciscala) monilifera (Melleville, 1843) (Pacaud herein).

perplexa , Scalaria – Deshayes, 1863: 60-61, pl. 8, fig. 1, ex Pease ms. — Type locality: La Réunion. — Type age: Recent. — Current status: synonym of Gyroscala lamellosa (Lamarck, 1822) [ Scalaria ] (L. G. Brown & Neville 2015: 121, following advice from Kilburn).

primula , Scalaria – Deshayes, 1861-DescrCoqFoss: 339, 11, pl. 11, figs 25-26. — Type locality: Mouy. — Type age: Eocene (Lutetian). — Type species (OD) of Cerithiscala Boury, 1887. — Current status: Tenuiscala (Cerithiscala) primula (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93; Pacaud 2008: 58).

propinqua , Scalaria – Deshayes, 1861-DescrCoqFoss: 345-346, 11, pl. 11, figs 31-32. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32411, syntype. Opalia (Pliciscala) gouldi [Deshayes, 1861] propinqua (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93) . — Current status: Opalia (Pliciscala) propinqua (Deshayes, 1861) ( Pacaud 2008: 58) .

sandbergeri , Scalaria – Deshayes, 1861-DescrCoqFoss: 335-336, 23, pl. 23, figs 5-6. — Type locality: Villepreux. — Type age: Oligocene (Rupelian). Preoccupies Scalaria rudis sandbergeri Sacco, 1891 . — Current status: Epitonium sandbergeri (Deshayes, 1861) ( Lozouet & Maestrati 2012b: 30) .

sculptata , Scalaria – Deshayes, 1861-DescrCoqFoss: 12, pl. 12, figs 8-9; 1865-DescrCoqFoss: 338-339. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian); 666 [error pro Scalaria cerithiformis Watelet, 1853 ]. This species is very similar to Foratiscala cerithiformis (Watelet, 1853) , with which Deshayes confused it; his taxon is now regarded as a full species. — Current status: Foratiscala sculptata (Deshayes, 1861) ( Le Renard & Pacaud 1995: 94) .

striatula , Scalaria – Deshayes, 1832-DescrCoqFoss: 198, 15, pl. 25, figs 6-8; 1843-HistNatAnim2: 81; 1843-HistNatAnim3: 531; 1861-DescrCoqFoss: 350. — Type locality: Châteaurouge. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32199, holotype. — Current status: Acirsa (Acirsella) striatula (Deshayes, 1861) ( Le Renard & Pacaud 1995: 94; Pacaud 2008: 58).

striatularis , Scalaria (Pyrgiscus) – Deshayes, 1861-DescrCoqFoss: 354, 12, pl. 12, fig. 16. — Type locality: Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Current status: synonym of Acirsa (Acirsella) striatula (Deshayes, 1832) (Pacaud herein).

subplicata , Scalaria – Deshayes, 1861-DescrCoqFoss: 351, 12, pl. 12, fig. 15. — Type locality: Châlons-sur-Vesle. — Type age: Paleocene (Thanetian). Acirsa (Plesioacirsa) subplicata (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93, 94). — Current status: Acirsa subplicata (Deshayes, 1861) ( Leroy et al. 2014: 25, pl. 21, figs 8-9.

tenuilamella , Scalaria – Deshayes, 1825-DescrCoqFoss: 13, pl. 22, figs 11-14; 1832: 195-196; 1843-HistNatAnim2: 82; 1843-HistNatAnim3: 531-532. 1861-DescrCoqFoss: 331-332. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32185, syntype. — Current status: Epitonium (Crisposcala) tenuilamella (Deshayes, 1825) ( Le Renard & Pacaud 1995: 92; Pacaud 2008: 58).

transversaria , Scalaria (Pyrgiscus) – Deshayes, 1861-DescrCoqFoss: 355, 12, pl. 12, fig. 20. — Type localities: Laon, Cuise-la-Motte & Laversine. — Type age: Eocene (Ypresian). — Current status: Acirsa (Acirsella) transversaria (Deshayes, 1861) ( Le Renard & Pacaud 1995: 94) .

turrella, Scalaria – Deshayes, 1861-DescrCoqFoss: 352, 11, pl. 11, figs 23-24. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32410, holotype. — Current status: synonym of Opalia (Dentiscala) plicata (Lamarck, 1804) (Pacaud herein).

variculosa , Scalaria –Deshayes, 1861-DescrCoqFoss: 349, 12, pl. 12, fig. 14. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Current status: Opalia (Crassiscala) variculosa (Deshayes, 1861) ( Le Renard & Pacaud 1995: 93; Pacaud 2008: 58).

vinsoni, Jantina [sic] – Deshayes, 1863: 94, 144 [corrected to Janthina ], pl. 11, figs 9, 11. — Type locality: La Réunion. — Type age: Recent. — Current status: synonym of Janthina exigua Lamarck, 1816 .

wardi, Scalaria – Deshayes, 1861-DescrCoqFoss: 352, 11, pl. 11, figs 17-19. — Type localities: Thiverval-Grignon, Parnes & Damery. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32408, 2 syntypes. — Current status: synonym of Opalia (Dentiscala) plicata (Lamarck, 1804) (Pacaud herein).

woodi , Scalaria –Deshayes, 1861-DescrCoqFoss: 339. — Type locality: England. — Type age: Coralline Crag, Pliocene. Amaea woodi (Deshayes, 1861) ( Marquet & Landau 2006: 34, fig. 8/4a-f). Not to be confused with Scalaria woodiana Nyst , in Omalius d’Halloy, 1862 & 1868, nor with Scalaria woodana Nyst , in Dewalque, 1868, both nomen nuda, now Scalaria hennei Nyst, 1871 . — Current status: synonym of Clathroscala cancellata ( Brocchi, 1814) [ Turbo ] ( Van Dingenen et al. 2016: 166-167, pl. 15, fig. 9).