Pupa reticulata ( Martin, 1884 )

Helwerda, Renate A., 2015, Acteonidae, Bullinidae and Ringiculidae (Gastropoda: Heterobranchia) from the Plio-Pleistocene of the Philippines, Zootaxa 3990 (2), pp. 197-220 : 204-206

publication ID	https://doi.org/ 10.11646/zootaxa.3990.2.2
publication LSID	lsid:zoobank.org:pub:BDF1BC11-4848-4858-99DC-0336A5FBEE2B
DOI	https://doi.org/10.5281/zenodo.6116513
persistent identifier	https://treatment.plazi.org/id/03FA878C-9517-FE43-FF26-E35D00BECBCD
treatment provided by	Plazi
scientific name	Pupa reticulata ( Martin, 1884 )
status

Treatment

Pupa reticulata ( Martin, 1884) View in CoL nov. comb.

( Figures 14–16 View FIGURE 14 – 18. 14 )

1884 Actaeon reticulatus nov. spec.—Martin: p. 43–44, pl. 4, fig. 42.

1982 Acteon reticulatus Martin—Shuto : p. 131, pl. 19, fig. 11.

2009 Acteon reticulatus Martin, 1884 View in CoL —Leloux & Wesselingh: p. 38, pl. 75, figs. 10–11.

Material. Tiep2 (6) RGM 961.789, Tiep3 (1) RGM 961.745, Tiep4 (1) RGM 961.790, Tiep5 (1) RGM 961.791.

Characterization. Shell tall; H 9.75 mm, W 4.42 mm; protoconch low, 1.1 whorl, DP 0.52 mm; nucleus small, DN 0.28 mm; teleoconch strongly sculpted with punctuated spiral grooves and many thin axial ribs forming a reticulate network; aperture narrow, expanded anteriorly; inner lip covered with thin callus, lamella present near columella; columella with pronounced double fold; umbilicus closed.

Remarks. The species has been assigned to Pupa on the basis of the strong double fold on the columella and the additional lamella on the inner lip above the columella. It is similar to some forms of P. solidula ( Linnaeus, 1758) in general shape and sculpture, but P. solidula ( Linnaeus, 1758) has weaker axial ribs, slightly narrower spiral grooves and a much weaker to obsolete lamella on the parietal wall. P. strigosa (Gould, 1859) and P. sekii Habe, 1958 have slightly narrower and fewer (20 and 23 respectively) spiral grooves, as well as weaker axial ribs. P. reticulata ( Martin, 1884) was originally described from the Burdigalian Miocene of Ngembak, Java, Indonesia, based on a single specimen (RGM 7302). Nine additional specimens from Late Burdigalian–Langhian Miocene were later found in Citalahab, Java, Indonesia (RGM 92176). Another specimen from the Miocene Paghumayan Formation of Negros Island, the Philippines, is described in Shuto (1982).