COLUMBELLIDAE SWAINSON, 1840

Kantor, Yuri I., Fedosov, Alexander E., Kosyan, Alisa R., Puillandre, Nicolas, Sorokin, Pavel A., Kano, Yasunori, Clark, Roger & Bouchet, Philippe, 2022, Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda), Zoological Journal of the Linnean Society 194, pp. 789-857 : 846-847

FAMILY COLUMBELLIDAE SWAINSON, 1840

= Anachidae Golikov & Starobogatov, 1972. Type genus: Anachis H.Adams & A.Adams, 1853 (TS Columbella scalarina G.B. Sowerby I, 1832 ; SD, Tate, 1868)

= Pyrenidae Suter, 1909. Type genus: Pyrene Röding, 1798 (TS Pyrene rhombiferum Röding, 1798 ; M)

SUBFAMILY COLUMBELLINAE SWAINSON, 1840

Type genus: Columbella Lamarck, 1799 (TS Voluta mercatoria Linnaeus, 1758; M).

SUBFAMILY ATILIINAE COSSMANN, 1901

Type genus: Atilia H.Adams & A.Adams, 1853 (TS Columbella suffusa G.B.Sowerby I, 1844 ; SD, Pace, 1902). Synonym: Anachis H.Adams & A.Adams, 1853 . (TS Columbella scalarina G.B.Sowerby I, 1832 ; SD, Tate, 1868).

Remarks: Columbellidae is a species-rich family, currently including 76 valid genera and 911 recent species (MolluscaBase, accessed 20.12.2020) and represented in our analysis by only seven species in six genera. It is recovered monophyletic with maximal support and sister to Cylleninae of Nassariidae (without support). Columbellidae possess a radula unique for Buccinoidea , with a rectangular acuspate central tooth and lateral teeth with narrow base and three to five cusps along the inner edge (see, e.g. Bandel, 1984). In Pseudanachis basedowi (Hedley, 1918) (not included in the analysis) the radula is markedly different in having multicuspidate central teeth and nearly subtriangular lateral teeth with more than ten small cusps ( Guralnick & deMaitenon, 1997).

UNNASSIGNED BUCCINOIDEA