Rubus malacocarpus Standl. & L.O.Williams

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Rubus malacocarpus Standl. & L.O.Williams ψ Arachnothryx buddleioides (Benth.) Planch.

Terrestrial shrubs. Distribution in PILA 2200–3200 m, low Algodoncillo (CR).

elevation oak forest, páramo, sabana natural grassland. Terrestrial shrubs or trees. Distribution in PILA Global distribution: Costa Rica, Panama. Records: B. 300–2700 m, low elevation mixed forest, mixed forest in Gamboa 1233 (INB, MO), E. Alfaro 3257 (INB), G. transition to low elevation forest, mixed forest in transition Davidse 24005 (CR, MO), G. Davidse 25185 (MO), G. to cloud forest, cloud forest, low elevation oak forest, Davidse 28910 (CR, MO), G. Davidse 29045 (MO), M. high elevation oak forest. Global distribution: Mexico Moraga 369 (INB). to Panama. Records: A.K. Monro 4905 (BM, MO), A.K. Monro 5683 (INB), A. Rodríguez 10972 (BM, INB, Rubus miser Liebm. PMA ), G. Herrera 6123 (MO). Regional conservation Terrestrial shrubs. Distribution in PILA 1600–3100 m, status: Near Threatened (NT).

mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, high elevation oak forest, páramo. Arachnothryx tayloriae (Lorence) Borhidi

Global distribution: Mexico to Panama. Records: A. Terrestrial shrubs or trees. Distribution in PILA 900–1500 Fernández 720 (CR, INB, MO), A. Fernández 912 (CR, m, mixed forest in transition to low elevation forest, mixed INB, MO), A. Soto 256 (INB), B.B. Klitgaard 786 (BM, forest in transition to cloud forest. Global distribution: INB, MO), C. Hamilton 775 (MO), D. Santamaría 7483 Endemic to Costa Rica. Records: A.K. Monro 5862 (BM, (PMA), D. Solano 4099 (BM, INB, MO, PMA), D. Solano INB, PMA), D. Solano 4830 (BM, INB, MO, PMA), G. 5286 (BM, INB, MO, PMA). Herrera 2656 (INB, MO), R. Aguilar 1116 (INB, MO).

Rubus urticifolius Poir. ψ Arcytophyllum lavarum K.Schum. ex Standl. Terrestrial shrubs. Distribution in PILA 500–2500 m, low Terrestrial shrubs. Distribution in PILA 800–3800 m, elevation mixed forest, mixed forest in transition to low mixed forest in transition to low elevation forest, mixed elevation forest, mixed forest in transition to cloud forest, forest in transition to cloud forest, cloud forest, low cloud forest, low elevation oak forest. Global distribution: elevation oak forest, páramo. Global distribution: Costa Mexico to Panama. Records: A. Rodríguez 11926 (BM, Rica, Panama. Records: A. Chacón 534 (INB, MO), B. INB, MO, PMA), A. Rodríguez 9769 (INB), A. Rodríguez Gamboa 1208 (CR, INB, MO), E. Alfaro 3275 (INB), 9850 (INB), A. Soto 1304 (INB), D. Santamaría 4428 E. Alfaro 3322 (INB), E. Alfaro 3397 (INB), G. Davidse (INB), D. Solano 2917 (INB), F. Quesada 1626 (INB, 25916 (CR, MO), G. Davidse 25951 (CR, MO), G. MO). Davidse 28826 (CR, MO), G. Davidse 29315 (CR, MO), G. Davidse 29342 (CR, MO), F. Quesada 1795 (INB),

RUBIACEAE W. Burger 8539 (MO). Global conservation status: Endangered (EN), B1ab(i,iii).

Alibertia atlantica (Dwyer) Delprete & C.H.Perss. Terrestrial trees. Distribution in PILA 0–1600 m, low Arcytophyllum muticum (Wedd.) Standl. elevation mixed forest, mixed forest in transition to low Terrestrial shrubs. Distribution in PILA 2700–3800 m, elevation forest, mixed forest in transition to cloud forest, high elevation oak forest, páramo. Global distribution: cloud forest. Global distribution: Nicaragua to Panama. Costa Rica to Ecuador, Venezuela. Records: E. Alfaro Records: G. Davidse 28514 (CR, MO), G. Herrera 3059 3327 (INB), G. Davidse 29297 (CR, MO). (MO). Bertiera guianensis Aubl.

Alibertia premontana C.M.Taylor Terrestrial shrubs. Distribution in PILA 0–800 m, low Terrestrial trees. Distribution in PILA 1000–1700 m, elevation mixed forest, mixed forest in transition to low mixed forest in transition to low elevation forest, mixed elevation forest. Global distribution: Mexico to Bolivia, forest in transition to cloud forest. Global distribution: Brazil. Records: G. Gallardo 60 (MO). Costa Rica, Panama. Records: D. Santamaría 5947 (BM, INB, PMA). Chimarrhis parviflora Standl. Pejibayillo, Manwood, Plátano amarillo, Ullama, Yema

Amaioua pedicellata Dwyer de huevo, Jagua amarillo (CR) . Terrestrial tree. Distribution in PILA 0–800 m, low Terrestrial herbs. Distribution in PILA 1000–1800 m, elevation mixed forest, mixed forest in transition to low mixed forest in transition to low elevation forest, mixed elevation forest. Global distribution: Honduras to Panama. forest in transition to cloud forest. Global distribution: Records: A. Rodríguez, Jul. 22, 2007 (Field obs.). Uses: Mexico to Brazil. Records: A. Rodríguez 11876 (BM, timber. INB, MO, PMA).

Chiococca pachyphylla Wernham Coccocypselum herbaceum P.Browne ex Aubl. Chiraquilla, Zorro (CR) . Terrestrial herbs. Distribution in PILA 0–500 m, low Terrestrial shrubs or trees. Distribution in PILA elevation mixed forest. Global distribution: Mexico to 200–3500 m, low elevation mixed forest, mixed forest in Panama. Records: A. Rodríguez 11411 (BM, INB, MO, transition to low elevation forest, mixed forest in transition PMA), G. Gallardo 77 (CR, INB, MO), G. Herrera 3244 to cloud forest, cloud forest, low elevation oak forest, (MO). páramo, sabana natural grassland. Global distribution: Mexico to Panama. Records: C. Hamilton 3873 (MO), E. Coccocypselum hirsutum Bartl. ex DC. Alfaro 4711 (INB). Huevos de indio (CR). Terrestrial herbs. Distribution in PILA 0–2000 m, low

Chione venosa (Sw.) Urb. elevation mixed forest, mixed forest in transition to low Terrestrial shrubs. Distribution in PILA 0–1800 m, low elevation forest, mixed forest in transition to cloud forest, elevation mixed forest, mixed forest in transition to low cloud forest. Global distribution: Mexico, Guatemala, elevation forest, mixed forest in transition to cloud forest, Costa Rica to Argentina. Records: A. Chacón 690 (MO), A. cloud forest. Global distribution: Mexico to Peru, Antilles. Rodríguez 9872 (INB), A. Rodríguez 9881 (INB), A. Soto Records: J. H. Kirkbride Jr. 158 (MO), K. E. Blum 2651 1305 (INB), D. Santamaría 3758 (INB), D. Santamaría (MO). 4430 (INB), D. Solano 2865 (INB), E. Alfaro 4192 (INB), E. Alfaro 4198 (INB), F. Quesada 1698 (CR, INB).

Chomelia venulosa W.C.Burger & C.M.Taylor Terrestrial tree. Distribution in PILA 200–900 m, low Coccocypselum lanceolatum (Ruiz & Pav.) Pers. elevation mixed forest, mixed forest in transition to low Terrestrial herbs. Distribution in PILA 1100–1900 m, elevation forest. Global distribution: Global distribution mixed forest in transition to low elevation forest, mixed Costa Rica, Panama. Records : A. Rodríguez, Nov. 1, 2007 forest in transition to cloud forest. Global distribution: (Field obs.) . Guatemala, Costa Rica to Argentina. Records: A. Rodríguez 9875 ( INB), D. Santamaría 4457 ( INB), D .

ψ Cinchona pubescens Vahl Solano 2868 (INB), D. Solano 3202 (INB), E. Alfaro Agujilla, Quina, Palo de lagarto, Quinina, Bichá (CR). 2202 (INB), E. Alfaro 4186 (INB), E. Alfaro 4189 (INB), Terrestrial trees. Distribution in PILA 900–2100 m, L. González 1221 (INB). mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low Condaminea corymbosa (Ruiz & Pav.) DC. elevation oak forest, dense thicket. Global distribution: Terrestrial shrubs. Distribution in PILA 0–2500 m, low Costa Rica to Bolivia, Venezuela. Records: D. Solano elevation mixed forest, mixed forest in transition to low 2897 (INB), G. Davidse 28466 (CR, MO), L. Acosta elevation forest, mixed forest in transition to cloud forest, 2280 (INB), R. Villalobos 257 (CR, INB). Uses: timber. cloud forest, low elevation oak forest. Global distribution: Regional conservation status: Near Threatened (NT). Costa Rica to Bolivia. Records: A.K. Monro 4697 (BM, INB).

Coccocypselum condalia Pers. Huevos de indio (CR). ψ Cosmibuena valerioi (Standl.) C.M.Taylor Terrestrial herbs. Distribution in PILA 1200–1700 m, Terrestrial shrubs or trees. Distribution in PILA 800–2600 mixed forest in transition to low elevation forest, mixed m, mixed forest in transition to low elevation forest, forest in transition to cloud forest. Global distribution: mixed forest in transition to cloud forest, cloud forest, Mexico to Panama, Peru, Venezuela, Brazil, Bolivia, low elevation oak forest, high elevation oak forest. Global Lesser Antilles. Records: A. Rodríguez 9872 (INB), distribution: Costa Rica, Panama. Records: G. Herrera A. Rodríguez 9881 (INB,), A. Soto 1305 (INB), D. 2698 (INB, MO), G. Herrera 3334 (INB, MO). Global Santamaría 3758 (INB), D. Santamaría 4430 (INB), D. conservation status: Vulnerable (VU), B1ab(i,iii). Solano 2865 (INB), E. Alfaro 4192 (INB), E. Alfaro 4198 (INB), F. Quesada 1698 (CR, INB). Coussarea caroliana Standl. Terrestrial shrubs or trees. Distribution in PILA 500–1600

Coccocypselum cordifolium Nees & Mart. m, low elevation mixed forest, mixed forest in transition to low elevation forest. Global distribution: Costa Rica,

Panama. Records: A. Rodríguez 11475 (BM, INB, MO, D. Solano 4157 (BM, INB, MO, PMA). Regional PMA), A. Rodríguez 11629 (BM, INB, MO, PMA), D. conservation status: Near Threatened (NT). Santamaría 5911 (BM, INB, MO, PMA), D. Santamaría 5929 (BM, INB, MO, PMA). Elaeagia chiriquina C.M.Taylor Terrestrial shrubs or trees. Distribution in PILA 1300–

Coussarea latifolia Standl. 1800 m, mixed forest in transition to cloud forest. Global Terrestrial trees. Distribution in PILA 200–600 m, low distribution: Costa Rica, Panama. Records: A.K. Monro elevation mixed forest. Global distribution: Costa Rica to 5752 (INB). Peru. Records: G. Herrera 3126 (MO). Elaeagia myriantha (Standl.) C.M.Taylor & Hammel

Coussarea talamancana Standl. Terrestrial trees. Distribution in PILA 600–1000 m, low Terrestrial shrubs or trees. Distribution in PILA 0–800 elevation mixed forest, mixed forest in transition to low m, low elevation mixed forest, mixed forest in transition elevation forest. Global distribution: Costa Rica, Panama, to low elevation forest. Global distribution: Costa Rica, Colombia, Ecuador, Peru, Bolivia, Venezuela. Records: Panama. Records: C. Davidson 6719 (MO). A. Rodríguez, Feb. 21, 2008 (Field obs.).

Crusea coccinea DC. Faramea eurycarpa Standl. Terrestrial herbs. Distribution in PILA 1800–2600 m, Azulejo (CR). cloud forest, low elevation oak forest, high elevation Terrestrial shrubs or trees. Distribution in PILA oak forest, sabana natural grassland. Global distribution: 300–1900 m, low elevation mixed forest, mixed forest in Mexico to Panama. Records: E. Alfaro 3471 (INB), G. transition to low elevation forest, mixed forest in transition Davidse 26136 (CR, MO), G. Davidse 28604 (CR, MO), to cloud forest, secondary forest. Global distribution: J. F. Morales 5823 (INB), M. Moraga 374 (INB). Costa Rica to Peru. Records: D. Solano 4863 (BM, INB, MO, PMA).

Declieuxia fruticosa (Willd. ex Roem. & Schult.) Kuntze Terrestrial herbs to shrubs. Distribution in PILA 0–2000 Faramea glandulosa Poepp. & Endl. m, low elevation mixed forest, mixed forest in transition Terrestrial shrubs. Distribution in PILA 0–1700 m, low to low elevation forest, mixed forest in transition to elevation mixed forest, mixed forest in transition to cloud forest, cloud forest. Global distribution: Mexico, low elevation forest, mixed forest in transition to cloud El Salvador, Nicaragua to Venezuela, Suriname, Brazil, forest, cloud forest. Global distribution: Mexico to Brazil. Paraguay. Records: A. Rodríguez 10305 (INB), D. Solano Records: González 774 (INB, PMA), D. Santamaría 7370 2863 (INB), E. Alfaro 4205 (INB). (INB, PMA), D. Santamaría 7851 (INB, PMA), D. Solano 4675 (BM, INB, MO, PMA), D. Solano 4792 (BM, INB,

Deppea grandiflora Schltdl. MO, PMA ), D. Solano 5657 (INB, PMA). Terrestrial shrubs. Distribution in PILA 1600–3000 m, mixed forest in transition to cloud forest, cloud forest, Faramea multiflora A.Rich. ex DC. low elevation oak forest, high elevation oak forest. Global Terrestrial shrubs or trees. Distribution in PILA 0–1600 distribution: Mexico to Panama. Records: D. Santamaría m, low elevation mixed forest, mixed forest in transition 7146 (INB). to low elevation forest, mixed forest in transition to cloud forest, cloud forest, dense thicket. Global distribution:

Didymaea mexicana Hook.f. Nicaragua to Bolivia. Records: A.K. Monro 4761 (BM, Terrestrial herbs. Distribution in PILA 1400–3400 m, INB, MO), A.K. Monro 6326 (PMA), D. Santamaría 6490 mixed forest in transition to cloud forest, cloud forest, low (BM, INB, MO, PMA). elevation oak forest, páramo. Global distribution: Mexico to Panama. Records: Davidse 26065 (INB), Davidse Faramea occidentalis (L.) A.Rich. 26160 (INB), Davidse 28789 (INB), Herrera 3637 (INB), Cafecillo (CR), Benjamín, Huesito, Garrotillo, Jazmín A.K. Monro 4059 (INB). (P). Terrestrial shrubs.Distribution in PILA 0–1200m, low

ψ Elaeagia auriculata Hemsl. elevation mixed forest, mixed forest in transition to low Gomilla, Huesillo, Madroño (CR). elevation forest. Global distribution: Tropical America. Terrestrial trees. Distribution in PILA 1200–1900 m, Records: A. Rodríguez, Nov. 1, 2007 (Field obs.), A. mixed forest in transition to low elevation forest, mixed Rodríguez, Sept 29, 2007 (Field obs.), D. Santamaría, forest in transition to cloud forest, pasture, dense thicket. Oct. 24, 2007 (Field obs.). Global distribution: Nicaragua to Panama. Records: D. Santamaría 3818 (INB, MO), D. Solano 2898 (INB), Faramea ovalis Standl. Terrestrial shrubs or trees. Distribution in PILA 1000–

3000 m, mixed forest in transition to low elevation forest, elevation oak forest, páramo, sabana natural grassland. mixed forest in transition to cloud forest, cloud forest, low Global distribution: Mexico to Panama. Records: A.K. elevation oak forest, high elevation oak forest, pasture. Monro 4104 (BM), C. Hamilton 3914 (MO), E. Alfaro Global distribution: Costa Rica to Ecuador. Records: A. 3204 (INB), E. Alfaro 3396 (INB). Rodríguez 11540 (BM, INB, MO, PMA), D. Santamaría 6665 (BM, INB, MO, PMA), D. Santamaría 7211 (BM, Gonzalagunia ovatifolia (Donn.Sm.) B.L.Rob. INB, MO, PMA ), D. Santamaría 7260 (BM, INB, MO, Terrestrial shrubs. Distribution in PILA 0–1400 m, low PMA), D. Solano 5204 (BM, INB, MO, PMA), G. Herrera elevation mixed forest, mixed forest in transition to low 2836 (MO), N. Zamora 4096 (BM, INB, MO, PMA). elevation forest, mixed forest in transition to cloud forest. Global distribution: Nicaragua to Colombia. Records:

Faramea scalaris Standl. A.K. Monro 6239 (PMA), D. Solano 4550 (BM, INB, Terrestrial shrubs. Distribution in PILA 300–1800 m, low MO, PMA). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Gonzalagunia rosea Standl. Global distribution: Costa Rica, Panama. Records: Alfaro Terrestrial shrubs or trees. Distribution in PILA 0–3000 5498 (MO). m, low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to

Faramea suerrensis (Donn.Sm.) Donn.Sm. cloud forest, cloud forest, low elevation oak forest, high Terrestrial shrubs or trees. Distribution in PILA 0–1700 elevation oak forest. Global distribution: Costa Rica to m, low elevation mixed forest, mixed forest in transition Colombia. Records: A.K. Monro 4220 (BM, INB, MO), A. to low elevation forest, mixed forest in transition to Soto 1246 (INB), C. Hamilton 3609 (MO), C. Hamilton cloud forest, cloud forest, pasture. Global distribution: 3640 (MO), D. Solano 5053 (BM, INB, MO, PMA), Nicaragua to Ecuador. Records: A. Jiménez 2818 (MO), E. Alfaro 743 (INB), E. Fletes 182 (INB), E. Mora 286 A. Rodríguez 11634 (BM, INB, MO, PMA), A. Rodríguez (CR, INB), G. Davidse 28469 (CR, MO), L. Acosta 2390 11664 (BM, INB, MO, PMA), D. Solano 4675 (BM, INB, (INB), M. Alfaro 144 (INB). Uses: ornamental. MO, PMA), D. Solano 4792 (BM, INB, MO, PMA). Gonzalagunia stenostachya (Standl.) W.C.Burger

Faramea uniflora Dwyer & M.V.Hayden Terrestrial shrubs. Distribution in PILA 400–2200 m, low Terrestrial shrubs. Distribution in PILA 1200– 1200 m, elevation mixed forest, mixed forest in transition to low mixed forest in transition to low elevation forest. Global elevation forest, mixed forest in transition to cloud forest, distribution: Costa Rica to Peru. Records: D. Santamaría cloud forest, low elevation oak forest. Global distribution: 6740 (BM, INB, MO, PMA), D. Solano 4120 (BM, INB, Costa Rica, Panama. Records: A. Rodríguez 11451 (BM, MO, PMA), D. Solano 4678 (BM, INB, MO, PMA), G. INB, MO, PMA), D. Santamaría 6582 (BM, INB, MO, Herrera 5373 (INB, MO). PMA), D. Solano 4688 (BM, INB, MO, PMA), D. Solano 4855 (BM, INB, MO, PMA), G. Herrera 5325 (CR, INB,

Galium aschenbornii Schauer MO ). Terrestrial herbs. Distribution in PILA 1400–3400 m, mixed forest in transition to cloud forest, cloud forest, low ψ Guettarda crispiflora Vahl elevation oak forest, páramo. Global distribution: Mexico Cantarillo,Mosqueta de montaña (CR) , Salvia de montaña, to El Salvador, Costa Rica, Panama, Ecuador. Records: E. Salvia de monte (P). Alfaro 4615 (INB). Terrestrial trees. Distribution in PILA 0–2700 m, low elevation mixed forest, mixed forest in transition to low

Galium hypocarpium (L.) Endl. ex Griseb. elevation forest, mixed forest in transition to cloud forest, Terrestrial herbs. Distribution in PILA 1000–3600 m, cloud forest, low elevation oak forest, high elevation oak mixed forest in transition to low elevation forest, mixed forest, pasture. Global distribution: Belize to Venezuela, forest in transition to low elevation forest, páramo. Global Bolivia, Greater Antilles, Lesser Antilles. Records: L. distribution: Mexico to Brazil, Chile. Records: A. Soto 281 Martínez 425 (PMA). Regional conservation status: (INB), D. Santamaría 3491 (INB), D. Santamaría 3786 Vulnerable (VU), A2c. (INB), D. Solano 2852 (INB), D. Solano 5104 (BM, INB, MO, PMA), E. Alfaro 3305 (INB), E. Alfaro 3728 (INB), Hamelia macrantha Little E. Alfaro 4190 (INB), G. Herrera 3704 (INB, MO), J. A. Terrestrial shrubs or trees. Distribution in PILA 0–1400 Duke 13699A (MO), F. Quesada 1587 (INB). m, low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to

Galium orizabense Hemsl. cloud forest. Global distribution: Costa Rica to Ecuador. Terrestrial herbs. Distribution in PILA 1300–3400 m, Records: A. Rodríguez 11284 (BM, INB, MO, PMA), A. mixed forest in transition to cloud forest, cloud forest, low Rodríguez 11452 (BM, INB, MO, PMA), D. Santamaría

6318 (BM, INB, MO, PMA), D. Santamaría 6579 (BM, m, low elevation mixed forest, mixed forest in transition INB, MO, PMA), D. Santamaría 7676 (PMA). to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, high

Hillia grayumii C.M.Taylor elevation oak forest. Global distribution: Costa Rica, Epiphytic shrubs. Distribution in PILA 0–800 m, low Panama. Records: L. Acosta 2365 (INB), J. Bittner 1749 elevation mixed forest, mixed forest in transition to low (INB, MO), K. E. Blum 2643 (MO), K. E. Blum 2599 elevation forest. Global distribution: Endemic to Costa (MO), A. Fernández 647 (CR, INB, MO), B. Hammel 1404 Rica. Records: D. Solano 4100 (BM, INB, MO, PMA). (MO), B. Hammel 1440 (MO), B. Hammel 5813 (MO), A. Rodríguez 10636 (BM, INB, MO, PMA), A. Rodríguez

Hillia loranthoides Standl. 10759 (BM, INB, MO, PMA), A. Rodríguez 11941 (BM, Epiphytic shrubs. Distribution in PILA 300–1900 m, low INB, MO, PMA), D. Santamaría 5728 (BM, INB, MO, elevation mixed forest, mixed forest in transition to low PMA), D. Solano 3992 (BM, INB, MO, PMA), D. Solano elevation forest, mixed forest in transition to cloud forest. 5051 (BM, INB, MO, PMA), D. Solano 5184 (BM, INB, Global distribution: Mexico to Guatemala., Costa Rica. MO, PMA), D. Solano 5195 (BM, INB, MO, PMA), N. Panama. Records: G. Herrera 6195 (CR, INB, MO). Zamora 4104 (BM, INB, MO, PMA), N. Zamora 4154 (BM, INB, MO, PMA), N. Zamora 4165 (BM, INB, MO,

Hillia maxonii Standl. PMA ), N. Zamora 4198 (BM, INB, MO, PMA). Epiphytic shrubs. Distribution in PILA 1300–2100 m,

mixed forest in transition to cloud forest, cloud forest, Hoffmannia areolata Standl.

low elevation oak forest, pasture. Global distribution: Terrestrial herbs to shrubs. Distribution in PILA 1400– Nicaragua to Ecuador. Records: A. Rodríguez 10832 2100 m, mixed forest in transition to cloud forest, cloud (BM, INB, MO, PMA), D. Solano 4048 (BM, INB, MO, forest, low elevation oak forest. Global distribution: Costa PMA). Rica, Panama. Records: A. De Sedas 692 (PMA), A.K. Monro 5427 (BM, INB, MO, PMA), A. Rodríguez 10800

Hillia panamensis Standl. (BM, INB, MO, PMA) , A. Rodríguez 10892 (BM, INB, Epiphytic shrubs. Distribution in PILA 0–1700 m, low MO, PMA), D. Solano 3871 (BM, INB, MO, PMA), D. elevation mixed forest, mixed forest in transition to low Solano 3946 (BM, INB, MO, PMA), E. Alfaro 5539 (BM, elevation forest, mixed forest in transition to cloud forest, INB), G. Davidse 29134 (MO), G. Herrera 5465 (INB, cloud forest. Global distribution: Mexico to Guatemala, MO), J. Lezcano 280 (PMA).

Nicaragua to Panama. Records: G. Davidse 25503 (MO).

Hoffmannia asclepiadea Standl.

Hillia triflora (Oerst.) C.M.Taylor Terrestrial herbs to shrubs. Distribution in PILA 1100– Tres flores (CR). 2700 m, mixed forest in transition to low elevation forest, Epiphytic shrubs. Distribution in PILA 800–2100 m, mixed forest in transition to cloud forest, cloud forest, mixed forest in transition to low elevation forest, mixed low elevation oak forest, high elevation oak forest. Global forest in transition to cloud forest, cloud forest, low distribution: Costa Rica, Panama. Records: D. Solano elevation oak forest. Global distribution: Nicaragua to 3990 (BM, INB, MO, PMA), E. Alfaro 4670 (INB), G. Colombia. Records: D. Santamaría 7017 (BM, INB, MO, Davidse 23779 (CR, MO), F. Quesada 1935 (INB). PMA), F. Quesada 1685 (INB).

Hoffmannia bullata L.O.Williams

Hoffmannia amplexifolia Standl. Terrestrial herbs. Distribution in PILA 100–1900 m, low Terrestrial herbs or shrubs. Distribution in PILA 1100– elevation mixed forest, mixed forest in transition to low 2300 m, mixed forest in transition to low elevation forest, elevation forest, mixed forest in transition to cloud forest. mixed forest in transition to cloud forest, cloud forest, low Global distribution: Mexico to Panama. Records: A. elevation oak forest. Global distribution: Costa Rica to Rodríguez 11320 (BM, INB, MO, PMA), D. Santamaría Colombia. Records: A. Fernández 768 (CR, INB, MO), A. 6273 (BM, INB, MO, PMA).

Fernández 868 (CR, INB, MO), A.K. Monro 4330 (BM,

INB, MO), A.K. Monro 5433 (BM, INB, MO, PMA), A.K. Hoffmannia congesta (Oerst.) Dwyer

Monro 5582 (BM, INB, MO, PMA), D. Santamaría 6613 Terrestrial herbs or shrubs. Distribution in PILA 400–1800 (BM, INB, MO, PMA), D. Santamaría 6717 (BM, INB, m, low elevation mixed forest, mixed forest in transition MO, PMA), D. Solano 3866 (BM, INB, MO, PMA), D. to low elevation forest, mixed forest in transition to cloud Solano 4813 (BM, INB, MO, PMA), G. Davidse 29189 forest. Global distribution: Costa Rica, Panama. Records: (MO), R. Aguilar 1096 (INB, MO). A. Fernández 759 (CR, INB, MO), A. Fernández 901 (CR, INB, MO), A.K. Monro 5438 (BM, INB, MO, PMA), A.K.

Hoffmannia arborescens Donn.Sm. Monro 6250 (PMA), A. Rodríguez 10588 (BM, INB, MO, Terrestrial herbs to shrubs. Distribution in PILA 300–3000 PMA), A. Rodríguez 11288 (BM, INB, MO, PMA), A. Rodríguez 11648 (BM, INB, MO, PMA), D. Santamaría m, low elevation mixed forest, mixed forest in transition 5809 (BM, INB, MO, PMA), D. Santamaría 6282 (BM, to low elevation forest. Global distribution: Costa Rica, INB, MO, PMA), D. Santamaría 6821 (BM, INB, MO, Panama. Records: G. Herrera 3054 (MO).

PMA), D. Solano 4416 (BM, INB, MO, PMA), D. Solano

4531 (BM, INB, MO, PMA), D. Solano 4532 (BM, Hoffmannia limonensis C.M.Taylor

INB, MO, PMA), S. Bridgewater 4177 (BM, INB, MO, Terrestrial herbs to shrubs. Distribution in PILA 200–1500 PMA). m, low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud

Hoffmannia davidsoniae Standl. forest. Global distribution: Costa Rica, Panama. Records: Terrestrial herbs or shrubs. Distribution in PILA 1100– A. De Sedas 691 (PMA), A.K. Monro 4738 (BM, INB, 2300 m, mixed forest in transition to low elevation forest, MO), A.K. Monro 5873 (BM, INB, MO, PMA), Rodríguez mixed forest in transition to cloud forest, cloud forest, low 11065 (BM, INB, MO, PMA), Rodríguez 11512 (BM, elevation oak forest. Global distribution: Costa Rica to INB, MO, PMA), D. Santamaría 5811 (BM, INB, MO, Colombia. Records: A. Rodríguez 10736 (BM, INB, MO, PMA), D. Santamaría 5978 (BM, INB, MO, PMA). PMA), A. Rodríguez 11818 (BM, INB, MO, PMA), D.

Santamaría 5751 (BM, INB, MO, PMA), D. Santamaría Hoffmannia longipetiolata Pol.

7085 (BM, INB, MO, PMA), D. Solano 3960 (BM, INB, Terrestrial shrubs. Distribution in PILA 0–2000 m, low MO, PMA), D. Solano 4023 (BM, INB, MO, PMA), D. elevation mixed forest, mixed forest in transition to low Solano 5055 (BM, INB, MO, PMA), D. Solano 5156 elevation forest, mixed forest in transition to cloud forest, (BM, INB, MO, PMA), G. Davidse 29120 (CR, MO), G. cloud forest, dense thicket. Global distribution: Costa Davidse 29175 (CR, MO), G. Herrera 5892 (CR, INB, Rica to Peru. Records: A. Fernández 972 (CR, INB, MO), MO), J. Dwyer 8756 (MO), J. Dwyer 8789 (MO), K. E. A.K. Monro 5766 (BM, INB, MO, PMA), A. Rodríguez Blum 2623 (MO). 11270 (BM, INB, MO, PMA), A. Rodríguez 11505 (BM, INB, MO, PMA), D. Santamaría 6311 (BM, INB, MO,

Hoffmannia dotae Standl. PMA ), D. Santamaría 6566 (BM, INB, MO, PMA), D. Terrestrial shrubs. Distribution in PILA 1000–2200 m, Solano 4000 (BM, INB, MO, PMA), D. Solano 4422 mixed forest in transition to low elevation forest, mixed (BM, INB, MO, PMA), D. Solano 4536 (BM, INB, MO, forest in transition to cloud forest, cloud forest, low PMA), D. Solano 4655 (BM, INB, MO, PMA), D. Solano elevation oak forest, dense thicket. Global distribution: 4735 (BM, INB, MO, PMA), E. Alfaro 4202 (INB), G. Costa Rica, Panama. Records: A. Fernández 779 (INB), Davidse 28372 (CR, MO), G. Herrera 3106 (INB, MO), A. Rodríguez 11827 (BM, INB, MO, PMA), A. Rodríguez M. Alfaro 139 (INB).

11923 (BM, INB, MO, PMA), A. Rodríguez 11959 (BM,

INB, MO, PMA), A. Soto 1237 (INB), D. Santamaría ε-PILA Hoffmannia manussatani Dwyer

7050 (BM, INB, MO, PMA), D. Santamaría 7091 (BM, Terrestrial herbs to shrubs. Distribution in PILA 1600– INB, MO, PMA), D. Santamaría 7278 (BM, INB, MO, 1900 m, mixed forest in transition to cloud forest. Global PMA), D. Solano 2909 (INB), D. Solano 5165 (BM, INB, distribution: Endemic to Panama. Records: B. Hammel MO, PMA), F. González 456 (INB), F. González 475 5811 (MO), S. Knapp 2122 (MO).

(INB), G. Davidse 23776 (CR, MO), F. Quesada 1650 This species has been documented from and adjacent (INB, MO), L. Acosta 2288 (INB), M. Alfaro 20 (INB), to the buffer zone of th epark but is still considered M. Moraga 427 (INB). endemic to PILA.