Aysenoides terricola, RAMÍREZ, 2003

RAMÍREZ, MARTÍN J., 2003, The Spider Subfamily Amaurobioidinae (Araneae, Anyphaenidae): A Phylogenetic Revision At The Generic Level, Bulletin of the American Museum of Natural History 2003 (277), pp. 1-262 : 67-70

Aysenoides terricola , new species Figure 27

TYPES: Male holotype and female paratype from Chile, Región V, Valparaíso province, Cuesta Cavilolén, ca. 31°46̍S, 71°19̍W, in soil crevices, 6.XI.1988, P. Goloboff, E. Maury, C. Szumik, deposited in MACN­Ar 9808.

ETYMOLOGY: The specific name refers to the habitat where the types were collected.

DIAGNOSIS: Resembles A. colecole in body shape and genitalia, but can be distinguished by having curved elevations on the epigyne anterior of the spermathecae, and by a longer cusp of the paramedian apophysis on the primary conductor.

FEMALE (paratype, fig. 27A): Total length 4.55. Carapace length 2.03, width 1.08, wid­ er on leg II. Length of tibia/metatarsus: I, 1.02/0.91; II, 0.87/0.81; III, 0.58/0.66; IV, 1.00/0.77. Palpal tarsus length 0.41. Chelicerae with three teeth on retromargin, basal one largest. Sternum length 1.06, width 0.64. Spines: leg I, femur d 1–1–1, p d1ap; tibia v 2­ 2­ 0­ 2 (the x­p1­ x displaced prolaterally); metatarsus v 2–2–0 (the x­p1­ x displaced prolaterally). II, femur = I; tibia v 2­ 2­ 0­ 2, p 0–1; metatarsus v 2­ r1–0. III, femur = I; tibia v r1ap or 0­ p1­ r1, p and r d1–1; metatarsus v 2–0–2 or r1–0–2, p and r 0­ d1–1, d 2ap. IV, femur d 1–1–1; tibia v p1­ p1–2; metatarsus v 2­ p1–2, p and r 1ap, d r1ap. Leg III directed forward. Femora I–III narrow, IV strong. Abdomen (slightly bowed ventrally) length 2.50, width 1.07. Color: carapace grayish, legs pale gray with brownish violet spots, I and II darker. Leg coxae pale, endites, labium, and sternum dark. Abdomen brownish violet with yellow dorsal pattern, venter brownish violet uniform. Epigyne (fig. 27D, E): lateral lobes anteriorly curved, widely separate, closer posteriorly, limiting T­shaped median field. Limit between lateral lobes and median field unclear close to copulatory openings. Spermathecae spherical, copulatory ducts heavily sclerotized.

MALE (holotype): Total length 4.00. Carapace length 1.87, width 1.07. Length of tibia/metatarsus: I, 1.63/1.47; II, 1.13/1.17; III, 0.73/0.87; IV, 1.23/1.03. Chelicerae smaller than those of female. Sternum length 0.98, width 0.64. Spines as in female, except: leg I, tibia v 2–2–2, p 1­ 0­ 1­ 0. II, tibia p 1­ d1­ 1­ 0; metatarsus v 2–2–0, p d1. III, tibia v 0­ p1­ r1; metatarsus v 2–2–2 or 2­ r1–2. IV, femur r d1ap; tibia v p1–2–2 or p1­ p1–2, r 1­ d1­ 1­ 0; metatarsus p 0­ d1–1, r d1­ d1–1. Abdomen length 2.13, width 0.97, spiracle– epigastrium 1.10, spiracle–spinnerets 0.11. Color as in female. Palp (fig. 27B, C): tibia width/length 0.58, RTA long, sharp. Cymbium relatively small, globose, cymbial conductor wide. Tegulum basal. Sperm duct with two conspicuous loops at dorsal anterior margin, one at base of secondary conductor. Embolus with basal process long, narrow. Median apophysis retrolateral, long, sinuous. Primary conductor with basal portion hyaline, without canal, not fitted to embolus; apical portion long, heavily sclerotized, with long canal, connected to basal portion by thin translucent vertical lamina. Secondary conductor triangular, with acute apex and membranous ventral area. Paramedian apophysis apparently with two cusps, one retrolateral, triangular, flat, another ventral, on primary conductor, long, narrow.

VARIABILITY: Male spines: III, tibia v 0– 2–2.

NATURAL HISTORY: The types were collected in crevices on dry soil, in a steep ravine (Pablo Goloboff, personal commun.).

DISTRIBUTION: Central Chile, in Elqui and Valparaíso provinces.

OTHER MATERIAL EXAMINED: CHILE: Región IV (Coquimbo): Elqui : 20 km N La Serena (Rt. 5 km 491), 120 m, 7.X.1992, N. Platnick, P. Goloboff, K. Catley , 13 ( AMNH). Región V (Valparaíso): Valparaíso: Same data as types, 3 immatures ( MACN­Ar) ; Viña del Mar , I.1979, A. Tobar, 13 3♀ 4 immatures ( AMNH) .

Figures

Fig. 27. Aysenoides terricola, n. sp., holotype and paratype A. Female, dorsal view. B. Male palp, ventral view. C. Same, retrolateral view. D. Epigyne, ventral view. E. Same, cleared. Scale bars = A, 1 mm; B–E, 0.2 mm. (C1 = primary conductor; MA = median apophysis; PMA = paramedian apophysis.)