Bacopa cladostyla Eskuche (1989: 11)

Sosa, María De Las Mercedes, Moroni, Pablo & O’Leary, Nataly, 2018, A taxonomic revision of the genus Bacopa (Gratioleae, Plantaginaceae) in Argentina, Phytotaxa 336 (1), pp. 1-27 : 11-14

publication ID

https://doi.org/ 10.11646/phytotaxa.336.1.1

persistent identifier

https://treatment.plazi.org/id/A76087F4-E54C-FF97-98F7-F9DFFD7B14AF

treatment provided by

Felipe

scientific name

Bacopa cladostyla Eskuche (1989: 11)
status

 

7. Bacopa repens (Swartz) Wettstein (1895: 76) View in CoL . Fig. 7 View FIGURE 7 .

Gratiola repens Sw. (1788: 14) View in CoL Herpestis repens (Sw.) Schlechtendal & Chamisso (1830: 107) View in CoL Herpestis obovata Poeppig. ex Chamisso & Schlechtendal (1830: 107) View in CoL pro. syn. ≡ Macuillamia repens (Sw.) Pennell (1923: 9) View in CoL .

Type:— JAMAICA. Jun. 1786, O. P. Swartz s.n. (lectotype S 04-341! [designated here], isolectotypes BM 000953377!, LINN-HS No. 48-3!, LD 1741638!, S-R-2471!, S-R-2470!).

= Macuillamia limosa Pennell (1920: 158) View in CoL Bacopa limosa (Pennell) Standley (1925: 460) View in CoL .

Type:— COLOMBIA. Cundinamarca: Melgar, open pool in clayey loam, alt. 400 – 500 m, F.W. Pennell 2927 (holotype NY 00090653, isotypes CAS 0003270!, PH 00016463!, US 00122518!).

= Bacopa cladostyla Eskuche (1989: 11) View in CoL , syn. nov.

Type:— PARAGUAY. San Bernardino, campo estero Ciervo-cuá, Feb 1915, T. Rojas 1043 (holotype LIL 222.457!, isotypes LIL 70.170 ex Hb. Gray!).

Herbs, annual, 3–5 cm tall, prostrate, ascending stems, strigose. Leaves opposite, sessile, blade ovate elliptic, 6–8 × 5–7 mm, apex rounded, base rounded, margin entire, adaxial surface glabrous, abaxial surface sparsely pubescent, more densely on the veins. Flowers axillary, solitary, pedicels 1–2(–5) mm long, villose; bracteoles absent. Calyx 5- lobed, the external dorsal lobe wide ovate, 2–4 × 2–2.8 mm, apex rounded, base cordate, the two lateral lobes ovate, 2–3 × 1.3–2 mm, apex obtuse, base cordate, strigose towards the apex and in the central vein, the two internal lobes linear, 2–4 × 0.4–0.6 mm, apex acute, hispid on margins and veins. Corolla white, tube 2.5–3 mm long, puberulous; stamens 2 plus 1 or 2 staminodes; ovary glabrous, style deeply bifid, stigmatic apex hairy globose. Capsule ovoid, apex acute, 2.8–3 × 2.2–2.5 mm.

Distribution and habitat:— Bacopa repens is found growing from the United States of America to Argentina, in the northeastern provinces of Corrientes, Formosa and Santa Fe. It is also frequent in China ( Deyuan et al. 1998). It grows between sea level and 500 m elev.

Nomeclatural remarks:— Swartz (1788) described Gratiola repens from material collected in Jamaica.According to Christenhusz (2011) the holotype is at S, where most of Swartz’s collections from the West Indies are housed ( Stafleu & Cowan 1986, Pupulin 2009). However, three sheets that can be undoubtedly attributed to Swartz were located there. Souza & Giulietti (2009: 54) lectotypified this name on a Bertero specimen from Jamaica although this element cannot be considered original material for the name since Swartz based the Prodromus on his own collections and those in the Banksian herbarium ( Nicolson & Jarvis 1990). Therefore, Souza & Giulietti’s designation is not in conformity with Art. 9.12 and must be superseded. Among the original material at S there is a specimen annotated in Swartz’s handwriting: “ Gratiola repens 1786 Jamaica ”. Thus, this specimen is here selected as lectotype of the name.

Taxonomic remarks:—The analysis of the type material of B. cladostyla , together with the description in the protologue evidences that it is a synonym of B. repens .

The presence of two stamens, instead of four, differentiates B. repens from the rest of the Bacopa that grow in Argentina. See remarks under B. australis .

Selected specimens examined:— Corrientes: Itatí, arrocera Repeczki, 28 Feb 1977, Ahumada et al. 891 (CTES). Formosa: Arrocera Sur. Establecimiento Negra-í, RN 11, km 1251, 20 Jan 2000, Iriart 577-8b (CTESN). Santa Fe: San Javier, 8 Feb 2000, Iriart 604-14ª (CTESN).

Kingdom

Plantae

Phylum

Tracheophyta

Class

Magnoliopsida

Order

Lamiales

Family

Plantaginaceae

Genus

Bacopa

Loc

Bacopa cladostyla Eskuche (1989: 11)

Sosa, María De Las Mercedes, Moroni, Pablo & O’Leary, Nataly 2018
2018
Loc

Bacopa cladostyla

Eskuche, U. 1989: )
1989
Loc

Macuillamia limosa

Standley, P. C. 1925: )
Pennell, F. W. 1920: )
1920
Loc

Gratiola repens

Pennell, F. W. 1923: )
Schlechtendal, D. & Chamisso, A. 1830: )
Schlechtendal, D. & Chamisso, A. 1830: )
1830
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