Bernaya squiresi, GROVES.

Groves, Lindsey & Squires, Richard L., 2023, Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover, PaleoBios 40 (10), pp. 1-52 : 15

BERNAYA SQUIRESI GROVES. 2011

FIGS. 5G–I

Bernaya (Bernaya) n. sp. Groves, 1997. p. 7 [as B. (B.) n. sp.].

Bernaya (Bernaya) squiresi Groves, 2011 . p. 46, 47, figs. 1, 2.

Bernaya squiresi Groves. Lorenz, 2017 . p. 208. Lorenz, 2018. p. 666, pl. 329, fig. 8.

Holotype and Type Locality— LACMIP 22341.1 , LACMIP Type 13644 ( Figs. 5G–I ), length 52.7 mm, width 40.0 mm, height 26.3 mm. LACMIP Locality 22341 [ex UCLA Locality 2341], Crescent Formation , southwest end of Discovery Bay, Jefferson County, southwestern Washington.

Occurrence— Middle lower Eocene (“Capay Stage”), southwest end of Discovery Bay , Jefferson County, southwestern Washington .

Etymology— Originally named for Richard L. Squires ( CSUN, Geological Sciences, professor emeritus).

Description— Shell of medium to large size, constricted on anterior end; maximum height near center of shell; maximum width slightly posterior of center; aperture wide, straight; dentition medium to coarse; labral (outer) lip with 24 teeth, columellar lip with 25 teeth; anterior terminal marginal callus (Groves 2011: p. 47).

Remarks— This species is known only from its holotype, which has moderately good preservation. Postburial crushing damaged the anterior portion of the aperture and also dorso-basally distorted the holotype. Perrilliat et al. (2003: pl. 1, figs. 5, 6) figured a specimen identified as Bernaya (Bernaya) obesa ( Deshayes, 1866) [hypotype IHN 5459] but is outwardly more similar to B. squiresi . Unfortunately, poor preservation prevents a positive identification. A poorly preserved outer lip fragment ( LACMIP 41599.2), which could be B. squiresi , from the Crescent Formation, LACMIP Locality 41599 (ex CSUN Locality 1599), Middle Fork Satsop River, Dry Lakes quadrangle, Mason County, Washington is noted. B. squiresi represents the northernmost occurrence of the genus in Cenozoic strata and the only representative of the genus described from Washington.

Figures

Figure 5 (previous page) A–C. Bernaya grovesi Squires and Demetrion, 1992, Bateque Formation (Eocene, “Ca- pay Stage”), Mesa La Salina, Baja California Sur, México. Basal (A), dorsal (B), and right (labral) (C) views, all 2.6x, length 17 mm, width 12 mm, height 7.7 mm, of holotype IGM 5172, LACMIP Locality 401220a (ex CSUN Locality 1220b). D–F. Bernaya kaylinae n. sp., Llajas Formation, (Eocene, “Domengine Stage”), Las Llajas Canyon, north side Simi Valley, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F) views, 3.8x, length 25.6 mm, width 17.3 mm, height 12.9 mm, of holotype, LACMIP 40374.80, LACMIP Type 14938 (LACMIP Locality 40374). G–I. Bernaya squiresi Groves, 2011, unnamed Eocene strata (“Capay Stage”), southwest end of Discovery Bay, Jefferson County, Washington. Basal (G), dorsal (H), and right (labral) side (I) views, 0.97x, length 52.7 mm, width 40 mm, height 26.3 mm, of holotype LACMIP 22341.1, LACMIP Type 13644, LACMIP Locality 22341. J–L. Bernaya sp., unnamed Paleocene strata (Thanetian Stage), near Lower Lake, Lake County, northern California. Basal (J), dorsal (K), and right (labral) side (L) views, 1.7x, length 33.8 mm, width 23 mm, height 18.3 mm, hypotype LACMIP 7047.72, LACMIP Type 14923 (LACMIP Locality 7047). M–O. Protocypraea? simiensis (Nelson, 1925), Santa Susana Formation (Thanetian Stage), Simi Hills, Ventura County, Paleocene, southern Cali- fornia. Basal (M), dorsal (N), and right (labral) side (O) views, 1.6x, length 32.3 mm, width 24.6 mm, height 17.3 mm, of holotype UCMP 30498, UCMP Locality 3818. P.Protocypraea? sp. 1, Oyster Bay Formation, Paleocene, Appian Way, Central Island District, Vancouver Island, British Columbia. Basal view, 1.8x, length 48.1 mm, width 31.4 mm, height 25.3 mm, of CDM 997.91.3, informal locality AP3.