Tetraodes Blanchard, 1853: 36

ROIG-JUÑENT, SERGIO, 2000, The Subtribes And Genera Of The Tribe Broscini (Coleoptera: Carabidae): Cladistic Analysis, Taxonomic Treatment, And Biogeographical Considerations, Bulletin of the American Museum of Natural History 2000 (1), pp. 1-90 : 48-49

Tetraodes Blanchard, 1853: 36 (type species Tetraodes laevis Blanchard 1853 by monotypy, = Cardiophthalmus clivinoides Curtis ).

TYPE SPECIES: (by original designation): Molops rivalis Germar, 1824 .

DIAGNOSIS: Barypus is the only broscine genus that has a distinctly developed basal keel on median lobe.

DESCRIPTION: External morphology (figs. 206–207, 212): Head with a single supraorbital puncture on each side, each bearing one seta; without temporal ridge; eyes emarginates; antenna with antennomeres 2–3 with a group of setae in the lateroposterior side, 4 setose on apical half, remaining pubescent all along the surface. Mandibles each with scrobal setae. Maxillar eustipes with three basal setae. Labium with two to four setae on the submentum; tooth of mentum bifid; prementum with glossal sclerite bisetose apically, ventral surface markedly keeled longitudinally. Pronotum not constricted apically, with two to six setae each side; prosternum with apical setae. Elytron with parascutellar striole isolated from apical portion of stria 1; parascutellar setae absent; rows of setiferous punctures in the 1st, 3rd, 5th, and 7th intervals present or absent; lateral umbilical series with 6–51 setae. Abdominal sterna 5–7 with a transverse groove. Foreleg trochanter with one or two setae; foretarsomeres 1–2 with or without adhesive vestiture, middle tarsomeres without it.

Male genitalia (figs. 208–210, 213–217): Sternum 9 with sclerotized ring complete with lateral margins narrow (figs. 213–214). Median lobe with dorsal surface sclerotized on basal half (fig. 210) and basal orifice completely closed dorsally, with basal keel distinctly developed (fig. 210, 217, bk). Internal sac without sclerites X and Y and apical plate (fig. 210). Left paramere asetose (figs. 208, 215); right paramere setose (figs. 209, 216).

Female genital tract (figs. 211, 218): Rami of gonocoxite 9 long. Gonopod 9 varies in size, with or without nematiform setae on subapical setose organ. Spermatheca short and broad (fig. 211, sp), joined in a common duct before entering the bursa copulatrix with median oviduct (fig. 218); helminthoid sclerite absent; bursa copulatrix and spermatheca without accessory glands.

HABITAT: This genus occurs in grassland areas of southern South America. The species inhabit Patagonian steppes, pampas of Uruguay and Argentina, grasslands of the Sierras of Córdoba, San Luis, and Salta provinces, in Argentina, and the bushy central regions of Chile.

GEOGRAPHICAL DISTRIBUTION: Barypus extends south of latitude 25° in Argentina, Uruguay, and central Chile.

MATERIAL EXAMINED: Barypus (A.) bonvouloiri (MLPA) ; B. (A.) parallelus (CIUC, MCZ) ; B. (B.) comechingonensis ( CM, IADIZA, MLPA); B. (B.) pulchellus (CUIC, MLPA) ; B. (B.) rivalis (CUIC, MLPA) ; B. (C.) clivinoides (CUIC, IADIZA, MCZ, MLPA) , B. (C.) chubutensis ( IADIZA); B. (C.) longitarsis (IADIZA, MLPA) ; and B. (C.) mendozensis (IADIZA) .

REMARKS: The systematics of the 22 species of Barypus has been recently elucidated by Roig-Juñent and Cicchino (1989) and Roig-Juñent (1992a, 1992b) who recognize the three subgenera proposed by Putzeys (1868) and conducted a cladistic analysis of their species (Roig-Juñent, 1995b).