Cinetus breviscapus, Brazidec & Vilhelmsen, 2022

Brazidec, Manuel & Vilhelmsen, Lars, 2022, New species of belytine and diapriine wasps (Hymenoptera: Diapriidae) from Eocene Baltic amber, European Journal of Taxonomy 813, pp. 57-86 : 62-64

Cinetus breviscapus sp. nov.

urn:lsid:zoobank.org:act:CCD7DD88-2504-4697-9B70-80EC819B1C24

Figs 1B , 2E–F , Table 1

Diagnosis

Head with sparse long hairs; antenna with uniform pilosity; scape less than five times as long as wide; pedicel rounded, as long as wide; flagellomeres cylindrical, elongate, longer than wide; F1 1.2 times as long as scape, deeply emarginated ( Fig. 2F ); anterior scutellar pits bean-shaped; propodeum with strong plicae; median propodeal keel simple; marginal vein longer than its distance from basal vein and as long as radial cell; postmarginal vein present on half radial cell; Rs directed toward base of wing; hind wing with basal cell open ( Fig. 1B ); hind tibia and tarsomeres with long recumbent hairs; petiole 2.8 times as long as high, with longitudinal ribs; gaster globose; T2 covering two thirds of gaster; S2 longer than T2 ( Fig. 2E ).

Etymology

The specific epithet is to be treated as an adjective and is composed of ‘ brevi- ’, meaning ‘short’, and ‘- scapus ’, a reference to the small size of the scape.

Type material

Holotype NHMD-300622 , a complete male.

Locality and horizon

Baltic amber is considered to be of Bartonian–Priabonian age, ca 34–38 Ma.

Description

Male

BODY. BL = 3.61 mm. Head with sparse long hairs (HeL = 0.47 mm); eye oval, glabrous; antennae inserted on transverse shelf; 14 antennomeres with uniform pilosity; scape elongate, less than five times as long as wide; pedicel rounded, as long as wide; flagellomeres elongate, cylindrical; F1 1.2 times as long as scape, with a deep emargination covering almost half of segment; F2–F6 slightly shorter, of subequal length; F6–F12 decreasing in size but still much more longer than wide; F12 tapering at apex (antennomeres length, in mm, when possible: Sc-0.32; P-0.06; F1-0.39; F2-0.34; F3-0.32); mandibles of ordinary form, not forming a beak.

MESOSOMA. Shorter than metasoma (MsL = 1.21 mm); pronotum without epomia, only visible in lateral view; mesoscutum smooth; notauli deep, diverging at posterior terminations toward a point outside scutellum; anterior scutellar pit bean-shaped; scutellum without posterior scutellar pits; mesopleuron with epicnemial pit, epicnemial bridge and subalar bridge present; propodeum with prominent plicae; median propodeal keel simple. Fore wing hyaline, uniformly micropubescent (FwL= 3.58 mm); venation complete with C, Sc+R, basal vein, M+Cu, marginal vein, r-rs and Rs pigmented; marginal vein very long, longer than its distance from basal vein and as long as radial cell; postmarginal vein fading after middle of radial cell; Rs proximally directed toward base of wing; M inconspicuous towards apex; Cu short and nebulous; radial cell closed. Hind wing narrow, with C pigmented and basal cell opened (HwL = 2.23 mm). Legs slender, covered with setae; tibial spur formula 1-2-2; hind tibia and tarsomeres with long recumbent hairs.

METASOMA. Petiole elongate, almost three times as long as its middle height (PtL = 0.56 mm; PtW = 0.19 mm), with longitudinal ribs; gaster globose, oval, smooth (GL = 1.37 mm; GH = 0.71 mm); six segments identifiable; T2 longest, covering two thirds of gaster; other tergites reduced, decreasing in length until they become indistinguishable; S2 longest, longer than T2; following sternites very short; pygidium and hypopygium at least as long as previous segments; genitalia internalized.

Female

Unknown.

Comments

Using Nixon’s (1957) key, Cinetus breviscapus sp. nov. keys out to Cinetus Jurine, 1807 because of the following characters: mandibles of ordinary form, scutellum without a row of foveae posteriorly, marginal vein very long, notauli slightly divergent posteriorly. Following Nixon’s (1957) key to male species of Cinetus , Cinetus breviscapus keys out near C. aletes Nixon, 1957 because of its scape that is shorter than F1 but differs in having the first flagellomere less distinctly longer than the scape (ratio scape length/F1 length: 1.2 vs 1.43; Nixon 1957), the petiole longer (three times as long as wide vs “about two and a half times as long as wide”) and being bigger (body length ca 3.6 mm vs ca 2.2 mm). In Baltic amber, Cinetus breviscapus sp. nov. is considered as separate from Cinetus inclusus Maneval, 1938 and Cinetus balticus Szabó & Oehlke, 1986 because it clearly does not fit with their descriptions. The petiole of C. balticus is longer than that of C. breviscapus (ratio length/middle height: 4 vs 2.8) and the fore wing venation is different ( C. breviscapus has a longer marginal vein, a longer radial cell, the postmarginal vein does not extend beyond the radial cell and Rs is conspicuous for almost all its length). Cinetus inclusus has a shorter petiole (ratio length/middle height: 1.5) and the scape as long as pedicel, F1 and half of F2 combined, unlike C. breviscapus whose scape is shorter than flagellomere 1.

Figures

Fig. 1. Wing venation of the species described here. A. Belyta knudhoejgaardi sp. nov. B. Cinetus breviscapus sp. nov.C. Cinetus elongatus sp. nov.D. Pantoclis globosasp. nov. E. Pantolyta augustinusii sp. nov. F. Basalys villumi sp. nov. G. Doliopria baltica sp. nov. H. Spilomicrus succinalis sp. nov. Scale bars: A–E = 1 mm; F, H = 0.5 mm; G = 0.25 mm.

Fig. 2. A–D. Belyta knudhoejgaardi sp. nov. A–C. Holotype (NHMD-608408). A. habitus in lateral view. B. Mesosoma and petiole in lateral view. C. Petiole and gaster in dorsal view. D. Paratype NHMD-608400, habitus in lateral view. E–F. Cinetus breviscapus sp. nov., holotype (NHMD-300622). E. Habitus in lateral view. F. Head in lateral view. Abbreviations: fe = flagellomere emargination; mlr = mesopleural longitudinal ridge; mpk = median propodeal keel. Scale bars: A = 2 mm; B, F = 0.5 mm; C–E = 1 mm.