Clavatula ’ irisae

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 33-35

‘ Clavatula ’ irisae nov. sp.

Figs 12F 1 –F 3 , G 1 –G 3 , 13A 1 –A 3 , B 1 –B 3, 3K , 5 , 6

Pleurotoma interrupta Brocc. — Hörnes 1854: 339, pl. 36, figs 19a–c] [non ‘ Clavatula ’ interrupta ( Brocchi, 1814) ].

Clavatula (Clavatula) olgae (Hoernes R. & Auinger, 1891) — Mikuž 1998: 74, pl. 4, figs 3–4 [non ‘ Clavatula ’ olgae ( Hoernes & Auinger, 1891) ].

Type material. Holotype: NHMW 2021 /0126/0001, SL: 42.9 mm, MD: 18.3 mm, Gainfarn ( Austria), Figs 13A 1 – A 3, 3K . Paratypes: NHMW 2021 /0126/0002, SL: 39.2 mm , MD: 17.5 mm, Gainfarn ( Austria) , Figs 13B 1 –B 3 ; NHMW 2021 /0126/0003, SL: 43.7 mm , MD: 17.5 mm, Gainfarn ( Austria) , Figs 12F 1 –F 2 ; NHMW 1846 /0037/0307, Enzesfeld ( Austria) , SL: 36.3 mm, MD: 15.7 mm, illustrated in Hörnes (1854, pl. 36, figs 19a–c). Figs 12G 1 –G 3 . NHMW 1855 /0045/0487, 16 spec., Gainfarn ( Austria) , SMF 351874 /4, 351875/1, Gainfarn ( Austria) .

Type locality. Gainfarn ( Austria), Vienna Basin .

Type stratum. Silt sand of the Baden Formation.

Age. Middle Miocene, middle Badenian (late Langhian).

Etymology. In honor of Iris Feichtinger (NHMW, Vienna, Austria), in recognition of her contributions to Miocene ichthyology.

Diagnosis. ‘ Clavatula ’ species of moderately large size, solid, stout fusiform shell with tripartite sculpture on early teleoconch whorls, later whorls with broad raised subsutural collar, narrow subsutural ramp, mid-whorl cord disappears about fifth whorl, suprasutural cord tuberculate, last whorl with strong but irregular tuberculate spiral sculpture, outer lip lirate within, siphonal canal moderately long.

Description. Shell moderately large, solid, stout fusiform with high spire; apical angle ~37–40°. Protoconch not preserved. Teleoconch of ten whorls. Early teleoconch whorls flat-sided with tripartite sculpture; prominent, flat subsutural cord, suprasutural cord with close-set, prominent beads, separated by narrow interspace with finely beaded central cord. Later whorls with broad, raised subsutural collar, bearing subobsolete, widely spaced, open spinous tubercles; deeply concave mid-portion, fine beaded cord disappears. Suprasutural cord with prominent opisthocline tubercles. Entire shells surface covered by numerous secondary spiral cords also overriding tubercles. Suture narrowly incised, running below beads. Last whorl 60% of total height; subsutural collar broad, bearing low, open spines just below suture, subsutural ramp narrow, deeply concave. Shoulder prominent, tubercular, mid-whorl weakly convex, moderately angled at peribasal cord. Base strongly constricted. Siphonal fasciole weakly swollen, twisted. Sculpture below ramp of strengthened shoulder and peribasal cords, with irregular, flattened, weakly tubercular cords of primary to tertiary strength. Aperture moderately wide, pyriform. Outer lip not thickened, with about 8–12 lirae deep within. Anal sinus deep, moderately wide, asymmetrically V-shaped, with apex below subsutural collar. Siphonal canal moderately long, narrow, straight to slightly deflected to left, without notch at tip. Columella excavated in upper third, weakly twisted below, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming broad callus rim and narrow pseudumbilical chink.

Discussion. This species is reminiscent of ‘ Clavatula ’ sophiae ( Hoernes & Auinger, 1891) , but differs in its stouter outline and shorter siphonal canal. Moreover, ‘ C. ’ sophiae lacks a prominent central beaded spiral cord on early teleoconch whorls, and its subsutural spiral cord has beads on early whorls, but is smooth in ‘ C. ’ irisae. ‘ Clavatula ’ olgae ( Hoernes & Auinger, 1891) has a comparable profile of the last whorl with a strong shoulder angulation, but differs clearly in its much higher spire and slender outline. ‘ Clavatula ’ vitalisi Strausz, 1955 is reminiscent of ‘ C. ’ irisae in general shape, but is distinguished by its slightly wider apical angle and the cancellate sculpture on the suprasutural cord. ‘ Clavatula ’ hirmetzli Kovács & Vicián, 2021 has an even stouter outline, coarser sculpture on the base, and lacks lirae inside the aperture.

Paleoenvironment. The common occurrence at Gainfarn and Enzesfeld suggests a preference for inner neritic environments with sea grass meadows ( Zuschin et al. 2007).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Gainfarn, Enzesfeld ( Austria) (hoc opus); Krka Basin: Dolenja Brezovica in Gorenja Stara vas ( Slovenia) ( Mikuž 1998).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia).

FIGURE 12 A1–A3. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). B1–B3. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). C1–C2. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, holotype, PAL 2021.1.1., Letkés (Hungary). D1–D2. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). E. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, paratype, PAL 2021.2.1., Letkés (Hungary). F1–F3. ‘Clavatula’ irisae nov. sp., paratype, NHMW 2021/0126/0003, Gainfarn (Austria). G1–G3. ‘Clavatula’ irisae nov. sp., paratype, NHMW 1846/0037/0307, Enzesfeld (Austria).

FIGURE 13 A1–A3. ‘Clavatula’ irisae nov. sp., holotype, NHMW 2021/0126/0001, Gainfarn (Austria). B1–B3. ‘Clavatula’ irisae nov. sp., paratype, NHMW 2021/0126/0002, Gainfarn (Austria). C1–C3. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). D1–D3. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), paralectotype, NHMW 2021/0127/0001,Enzesfeld (Austria).E1–E3. ‘Clavatula’ sophiae (Hoernes &Auinger, 1891), NHMW 2021/0126/0004, Gainfarn (Austria), F1–F3. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), NHMW 2021/0126/0005, Gainfarn (Austria).