NERITIDAE Rafinesque, 1815

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 586-588

Family NERITIDAE Rafinesque, 1815

Cliton – Deshayes, 1830a-EncyMeth: 246. Incorrect subsequent spelling of Clithon Montfort, 1810 .

Tomostoma – Deshayes, 1824b: 187. — Type species (SD Fischer, 1885): Pilelolus neritoides Deshayes, 1824 . In erecting this genus, Deshayes noted that he originally proposed Pilelolus at a meeting, but then learned that G. B. Sowerby I, 1823, had established Pileolus J. Sowerby, 1823 (SD J. E. Gray, 1847), for P. plicatus G. B. Sowerby I, 1823, and other species, which Deshayes regarded belonging in this group. Tomostoma was thus proposed in synonymy. However, the ICZN Code Art. 11.6.1 provides that a name established in synonymy can become available if it was used as valid before 1961, as was this one. Symonds & Pacaud (2010: 55-56) discussed the history of these two genus-group names and determined that they are not confamilial – Pileolus J. Sowerby, 1823 , belongs in the Pileolidae . — Current status: Tomostoma is a valid genus in the Neritidae . Tomastoma G. B. Sowerby I, 1824, is a subsequent incorrect spelling ( Petit 2009: 196-197, note 48).

angistoma , Nerita – Deshayes, 1825-DescrCoqFoss: 12, pl. 29, figs 11-12; 1832: 159; 1832-EncyMeth: 622;1838-HistNatAnim2: 618-619; 1843-HistNatAnim3: 490; 1857-TraitElem: x, pl. 73 ter, figs 6-7, 10; 1864-DescrCoqFoss: 13, 66, pl. 66, figs 4-6; 1865-DescrCoqFoss: 666 [should have been Nerita angystoma , but this change would not be allowed]. — Type locality: Valmondois. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32173, holotype. — Current status: Otostoma angistoma (Deshayes, 1825) ( Le Renard & Pacaud 1995: 89) . Nerita angistoma Deshayes, 1825 ( Pacaud 2009a: 351).

auversiensis , Nerita – Deshayes, 1864-DescrCoqFoss: 15-16, 66, pl. 66, figs 12-14. — Type localities: Auvers-sur-Oise, Valmondois, Le Fayel & Yvors. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32667, syntype from Le Fayel. — Current status: Puperita (Heminerita) auversiensis (Deshayes, 1864) ( Le Renard & Pacaud 1995: 90) .

bicoronata , Nerita –Deshayes, 1864-DescrCoqFoss: 13-14, 66, pl. 66, figs 10-11. — Type localities: Noailles & Aizy-Jouy. — Type age: Paleocene-Eocene (Thanetian-Ypresian).— Type material:UCBL-EM 32666, lectotype from Noailles ( Pacaud 2009a). Otostoma bicoronata (Deshayes, 1864) ( Le Renard & Pacaud 1995: 89) . — Current status: synonym of Otostoma pouechi d’Archiac, 1859 ( Pacaud 2009a: 353).

brimonti , Nerita – Deshayes, 1864-DescrCoqFoss: 14-15, 66, pl. 66, figs 15-17. — Type locality: Brimont. — Type age: Paleocene (Thanetian). — Type material: UCBL-EM 32668, holotype. Nerita (Amphinerita) brimonti Deshayes, 1864 ( Le Renard & Pacaud 1995: 89). — Current status: Monsneritina brimonti (Deshayes, 1864) (Pacaud herein).

callosa , Neritina –Deshayes, 1835b: 156, pl. 19, figs 16-18;1838-HistNatAnim2: 591; 1843-HistNatAnim3: 480. — Type locality: Peloponnese Peninsula, Greece. — Type age: Recent. Theodoxus (Neritaea) varius callosus (Deshayes, 1835) ( Bank 2006: 52). — Current status: Theodoxus (Theodoxus) callosus (Deshayes, 1835) ( Eichhorst 2016: 905, 976, pl. 272, as “1833”).

consobrina , Neritina – Deshayes, 1832-DescrCoqFoss: 153, pl. 20, figs 5-6, ex Férussac ms. — Type localities: Épernay & Cumière. — Type age: Eocene (Ypresian). Incorrect subsequent spelling of Nerita sobrina Férussac, 1823 . Clithon (Vittoclithon) sobrinus (Férussac, 1823) ( Pacaud 2017c: 194-195) . Clithon (Pictoneritina) sobrinus (Férussac, 1823) ( Vrinat 2019: 51, pl. 10, figs 1-6, pl. 11, figs 1-6). — Current status: Clithon sobrinum (Férussac, 1823) (Pacaud herein).

costulata , Nerita – Deshayes, 1838-HistNatAnim2: 617;1843-HistNatAnim3: 489-490. Replacement name for Nerita costata J. de C. Sowerby, 1824 (J. de C. Sowerby 1824: 94, pl. 463, figs 5-6), “ non Chemnitz”, actually non Gmelin, 1791, nec Brocchi, 1814, nec Schumacher, 1817. — Type species (OD) of Cautochilus Gründel, 2004 . — Type locality: Great Britain. — Type age: Jurassic. Preoccupies Nerita costulata von dem Busch, 1844 . Lissochilus costulatus (Deshayes, 1838) (J.-C. Fischer & Weber 1997: 89, pl. 18, figs 14- 15). — Current status: Cautochilus costulatus (Deshayes, 1838) ( Gründel 2004: 33) .

crassula , Nerita – Deshayes, 1858: 553, nomen nudum.

duchasteli , Neritina – Deshayes, 1825-DescrCoqFoss: 11, pl. 17, figs 23-24; 1832: 154-155; 1838-HistNatAnim2: 595; 1843-HistNatAnim3: 482. 1864-DescrCoqFoss: 24-25. — Type locality: Versailles. — Type age: Oligocene (Rupelian). — Current status: Vitta duchasteli (Deshayes, 1825) ( Lozouet & Maestrati 2012a: 270, 273, fig. 175: 1-9).

dutemplei , Neritina – Deshayes, 1864-DescrCoqFoss: 21, 66, pl. 66, figs 21-24. — Type localities: Ay & Sinceny. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32670-32671, syntypes from Ay. Theodoxus (? Vittoclithon) dutemplei (Deshayes, 1864) ( Le Renard & Pacaud 1995: 90) . — Current status: Neritodryas dutemplei (Deshayes, 1864) ( Symonds & Pacaud 2010: 61-62, fig. 7; Vrinat 2019: 54, pl. 15, figs 1-5).

elegans , Neritina – Deshayes, 1825-DescrCoqFoss: pl. 19, figs 3-4; 1832: 154; 1838-HistNatAnim2: 595-596; 1843-HistNatAnim3: 482; 1864-DescrCoqFoss: 24. — Type locality: Houdan. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32169, syntype. Theodoxus (Vittoclithon) sublineolatus [d’Orbigny, 1850] elegans (Deshayes, 1825) ( Le Renard & Pacaud 1995: 90) . Clithon (Vittoclithon) elegans (Deshayes, 1825) ( Pacaud 2008: 45) . Clithon (Pictoneritina) elegans (Deshayes, 1832) ( Vrinat 2019: 48, pl. 2, fig. 1). — Current status: Clithon elegans (Deshayes, 1825) (Pacaud herein).

fulgurata, Neritina – Deshayes, 1863: 80-81, pl. 10, figs 1-2. — Type locality: La Réunion. — Type age: Recent. — Current status: synonym of Vittina gagates (Lamarck, 1822) [ Neritina ] ( Eichhorst 2016: 1088-1089, 1124, pl. 334).

graciosa, Nerita ; see: gratiosa, Nerita below.

granulosa , Nerita – Deshayes, 1825-DescrCoqFoss: 12, pl. 29, figs 13-14; 1832: 159-160; 1838-HistNatAnim2: 618; 1843-HistNatAnim3: 490; 1864-DescrCoqFoss: 16-17. — Type localities: Valmondois & Senlis. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32166, syntype from Valmondois. Nerita (Theliostyla) granulosa (Deshayes, 1825) ( Le Renard & Pacaud 1995: 89) . — Current status: Nerita (Theliostyla) granulosa (Deshayes, 1825) ( Le Renard & Pacaud 1995: 89) . — Current status: Monsneritina granulosa (Deshayes, 1832) (Pacaud herein).

gratiosa, Neritina – Deshayes, 1864-DescrCoqFoss: 19-20, 66, pl. 66, figs 27-29. In the body of the text on p. 19, the species was misspelled as N. “graciosa”. — Type localities: Brimont, Châlons-sur-Vesle, Gueux & Jonchery-sur-Vesle. — Type age: Paleocene (Thanetian). — Type material: UCBL-EM 32673, syntype from Jonchery-sur-Vesle. — Current status: synonym of Neritoplica subornata (d’Orbigny, 1850) [ Nerita ], replacement name for Neritina ornata Melleville, 1843 (Melleville 1843: 96-97 [50-51, 85], pl. 6, figs 9-10), non Nerita ornata G. B. Sowerby II, 1823 (Pacaud 2007: 30; Vrinat 2019: 55, pl. 2, fig. 5).

intermedia , Nerita – Deshayes, 1832b: 420-421, 439, 522, pl. 1, figs 6-7, non Grateloup, 1828. Nerita pileolus Récluz, 1850 , replacement name, but as “ non Neritina intermedia G. B. Sowerby I, 1833”, a non-homonym. — Type locality: Malabar, India; Bélanger. — Type age: Recent. — Type material: MNHN-IM-2000-32729, 4 syntypes. — Current status: synonym of Neripteron pileolus ( Récluz, 1850) [ Neretina ] ( Eichhorst 2016: 373, 411, pl. 90).

jaspidea, Neritina – Deshayes, 1864-DescrCoqFoss: 20, 65, pl. 65, figs 14-16. — Type localities: Brimont, Châlons-sur-Vesle & Gueux. — Type age: Paleocene (Thanetian). — Type material: UCBL-EM 33439-33457, 19 syntypes from Châlons-sur-Vesle.– Current status: synonym of Neritoplica vicina (Melleville, 1843) ( Glibert 1962a: 103) .

lamarckii , Neritina –Deshayes, 1838-HistNatAnim2: 578;1843-HistNatAnim3: 475-476. — Type locality: none given. — Type age: Recent. — Type material:MNHN-IM-2000-32731, 6 syntypes. — Current status: synonym of the Indo-Pacific Neripteron taitense (Lesson, 1831) [ Nerita ] ( Eichhorst 2016: 388-389, 421, pl. 100). lineolata , Neritina – Deshayes, 1825-DescrCoqFoss: 12, pl. 19, figs 7-8;1832: 152-153, non Lamarck, 1816; 1838-HistNatAnim2: 596; 1843-HistNatAnim3: 482. 1858: 553. 1864-DescrCoqFoss: 23-24. — Type locality: Houdan. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32171, syntypes. Synonym of Clithon inequidentata ( Récluz, 1850: 154) [April] [ Neritina ], replacement name. Nerita sublineolata d’Orbigny, 1850b (d’Orbigny 1850b: 346, no. 145) [November], unnecessary additional replacement name (Pacaud 2007: 30). — Current status: Clithon (Pictoneritina) inequidentata ( Récluz, 1850) ( Vrinat 2019: 48, pl. 2, figs 2-3). Clithon inequidentatum ( Récluz, 1850) (Pacaud herein).

modicella , Neritina –Deshayes, 1863: 79-80, pl. 10, figs 3-4. — Type locality: La Réunion. — Type age: Recent. — Current status: synonym of Vittina natalensis (Reeve, 1855) [ Nerita ] ( Eichhorst 2016: 1091-1092, 1126, pl. 336).

neritoides , Pileola [sic] – Deshayes, 1824b: 191-192, 461, as Pileolus , pl. 13, fig. 5a-c; 1825-DescrCoqFoss: 11, pl. 17, figs 17-18; 1832: 146-147; 1828-DictClass13: 568; Deshayes inGuérin-Méneville 1835: pl. 15, fig. 5, 5a; 1839-RegAnim: pl. 48 bis, fig. 1, 1a-c; Deshayes in Guérin-Méneville 1844: 30; 1864-DescrCoqFoss: 26; Guérin-Méneville & Deshayes 1868: [29], pl. 14, fig. 5, 5a, 5b. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Type species (SD Fischer, 1885) of Tomostoma Deshayes, 1824 (see under genus above). Pileolus (Tomostoma) altavillensis [Defrance, 1818; Crepidula ] neritoides (Deshayes, 1824) ( Le Renard & Pacaud 1995: 90, as “1825”). — Current status: Pileolus (Tomostoma) neritoides (Deshayes, 1824) ( Pacaud 2008: 45, as “1825”). Tomostoma altavillensis neritoides (Deshayes, 1824) ( Symonds & Pacaud 2010) .

nucleus , Neritina –Deshayes, 1832-DescrCoqFoss: 156-157, 15, pl. 25, figs 3-5; 1838-HistNatAnim2: 596-597;1843-HistNatAnim3: 482- 483; 1864-DescrCoqFoss: 23. — Type localities:Retheuil & Cuise-la-Motte. — Type age: Eocene (Ypresian). — Type material:UCBL-EM 32198, syntypes from Cuise-la-Motte. — Current status: Theodoxus (Vittoclithon) nucleus (Deshayes, 1832) ( Le Renard & Pacaud 1995: 90) . Clithon (Pictoneritina) nucleus (Deshayes, “1825”) ( Symonds & Pacaud 2010: 60). Clithon (Pictoneritina) nucleus (Deshayes, 1832) ( Vrinat 2019: 48-49, pl. 4, figs 1-6, pl. 5, figs 1-6). Clithon nucleus (Deshayes, 1832) (Pacaud herein).

passyana , Neritina – Deshayes, 1864-DescrCoqFoss: 24, 65, pl. 65, figs 11-13 [in text as figs 2-13];1865-DescrCoqFoss: 666 [correction noted].— Type localities: Montagny-en-Vexin, Le Fayel & Auvers-sur-Oise. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32660, syntypes from Montagny-en-Vexin. Theodoxus (Vittoclithon) passyanus (Deshayes, 1864) ( Le Renard & Pacaud 1995: 90) . Theodoxus (Pictoneritina) passyanus (Deshayes, 1864) ( Symonds, 2002: 3-6, pl. 1, figs 6-8, pl. 2, figs 9-14). — Current status: Clithon (Pictoneritina) passyanus (Deshayes, 1864) ( Symonds & Pacaud 2010: 65; Vrinat 2019: 50). Clithon passayanum (Deshayes, 1864) (Pacaud herein).

pentastoma, Nerita –Deshayes, 1864-DescrCoqFoss: 17-18, 66, pl. 66, figs 7-9. — Type localities:Mouchy-le-Châtel & Thiverval-Grignon.— Type age: Eocene (Lutetian). — Type material: UCBL-EM 32665, syntypes from Thiverval-Grignon. Nerita (Theliostyla) tricarinata [Lamarck, 1804] pentastoma Deshayes, 1864 ( Le Renard & Pacaud 1995: 89). Pseudodostia tricarinata pentastoma (Deshayes, 1864) ( Pacaud 2008: 45) . — Current status: Monsneritina tricarinata pentastoma (Deshayes, 1864) (Pacaud herein).

pyrenaica, Nerita –Deshayes, 1857-TraitElem: xi, pl. 73 ter, fig.17, non Nerita fluviatilis pyrenaica Moquin-Tandon, 1855 . — Type locality: Gensac. — Type age: Cretaceous (Maastrichtian). — Type material: UCBL-EM 32117, syntype. — Current status: synonym of Otostoma retzii (Nilsson, 1827) [ Natica ] (Pacaud herein).

saincenyensis , Neritina – Deshayes, 1864-DescrCoqFoss: 21-22, 66, pl. 66, figs 25-26. — Type locality: Sinceny. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32672, syntypes. Theodoxus saincenyensis (Deshayes, 1864) ( Le Renard & Pacaud 1995: 90) . Clithon (Pictoneritina) saincenyensis (Deshayes, 1832) ( Vrinat 2019: 50-51, pl. 9, figs 1-6, pl. 15, fig. 6). — Current status: Clithon saincenyense (Deshayes, 1864) (Pacaud herein).

sandwichensis, Neritina –Deshayes, 1838-HistNatAnim2: 579. —Type locality:Hawaiian Islands.— Type age: Recent. 1843-HistNatAnim3: 476. — Type material:MNHN-IM-2000-32762, 3 syntypes. — Current status: synonym of the endemic Neripteron (Dostia) cariosum (W. Wood, 1828) [ Nerita ] ( Pease 1871: 101; Kay 1979: 66, as Theodoxus ; Eichhorst 2016: 356-358, 411, pl. 90).

semilugubris , Nerita – Deshayes, 1864-DescrCoqFoss: 15, 66, pl. 66, figs 18-20. — Type localities: Brimont, Châlons-sur-Vesle & Jonchery-sur-Vesle. — Type age: Paleocene (Thanetian). — Type material: UCBL-EM 32669, syntypes. Nerita (Amphinerita) semilugubris Deshayes, 1864 ( Le Renard & Pacaud 1995: 89; Leroy et al. 2014: 23, pl. 16, figs 5-8). — Current status: Monsneritina semilugubris (Deshayes, 1864) (Pacaud herein).

zonaria, Neritina – Deshayes, 1832-DescrCoqFoss: 156, 15, pl. 25, figs 1-2; 1838-HistNatAnim2: 597; 1843-HistNatAnim3: 483; 1864-DescrCoqFoss: 23. — Type localities: Retheuil & Cuise-la-Motte. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 32197, syntype from Cuise-la-Motte. — Current status: Clithon (Vittoclithon) zonarius (Deshayes, 1832) ( Le Renard & Pacaud 1995: 90) . Clithon (Pictoneritina) zonarius (Deshayes, 1832) ( Vrinat 2019: 53-54, pl. 13, figs 1-6, pl. 14, figs 1-3). — Current status: Clithon zonarium (Deshayes, 1832) (Pacaud herein).

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danubialis , Neritina – Deshayes, 1838a: 65, pl. 5, figs 4-5. Sherborn (1925: 1784) listed this as a Deshayes species. However, it was made available by C. Pfeiffer (1828) and is now known as Theodoxus danubialis (C. Pfeiffer, 1828) .