Cunnolites polymorphus ( Goldfuss, 1826 )

Baron-Szabo, Rosemarie C., 2008, Dendrophylliina, Caryophylliina, Fungiina, Microsolenina, and Stylinina, Zootaxa 1952, pp. 1-244 : 173-176

Cunnolites polymorphus ( Goldfuss, 1826)

Text-Fig. 33

Text-Fig. 33 Cunnolites polymorphus ( Goldfuss, 1826) , as figured in Duncan (1880), as type material of Cyclolites vicaryi Duncan, 1880 , Paleocene of India. A, lateral view of corallum, upper surface, scale bar: 7 mm; B, cross view, close-up, scale bar: 1 mm.

v*1826 Fungia polymorpha nobis: Goldfuss, vol. 1, p. 48, pl. 14, figs 6 a–c, g–m.

v1826 Fungia undulata nobis: Goldfuss, vol. 1, p. 49, pl. 14, fig. 7.

v1826 Fungia discoidea nobis: Goldfuss, vol. 1, p. 50, pl. 14, fig. 7.

1834 Cyclolites discoidea, Goldf. : Blainville, p. 335.

1847 Cyclolites hemisphaerica n.: Michelin, p. 282, pl. 64, figs 2a–b.

1850 Funginella discoidea: d‘Orbigny , vol. 2, p. 202.

v1854 Cyclolites placenta m.: Reuss, p. 125, pl. 17, figs 4–6.

1851b Cyclolites discoidea: Milne Edwards & Haime , vol. 2, p. 202.

1851 Cunnolites semigloboso Michelin : Leymerie, p. 199, pl. 8, fig. 14.

1858–61 Cyclolites placenta: de Fromentel , p. 124.

1860 Cyclolites polymorpha: Milne Edwards , vol. 3, p. 44.

1863 Cyclolites polymorpha: de Fromentel , p. 336, pl. 59, pl. 60, fig. 1.

1863 Cyclolites discoidea: de Fromentel , p. 351, pl. 65, pl. 66, fig. 1, pl. 69, fig. 2.

1867 Cyclolites placenta Reuss : de Fromentel, p. 333.

1870 Cyclolites reussi: de Fromentel , p. 338, pl. 55, fig. 3.

1880 Cyclolites Ranikoti, Duncan : Duncan, p. 52, pl. 17, figs 5–6.

1880 Cyclolites vicaryi, Haime : Duncan, p. 53, pl. 17, figs 1–2.

1880 Cyclolites regularis: Leymerie , p. 774.

1881 Fungia elliptica Lamarck : Quenstedt, p. 861, pl. 177, fig. 1.

1881 Fungia undulata rotunda: Quenstedt , p. 862, pl. 177, figs 2–3.

1881 Fungia dispar: Quenstedt , p. 862, pl. 177, fig. 4 (non fig. 24).

1881 Fungia polymorpha Goldfuss : Quenstedt, p. 863, pl. 177, fig. 7.

1881 Fungia discoidea: Quenstedt , p. 865, pl. 177, fig. 14.

1881 Fungia patellata: Quenstedt , p. 871, pl. 177, fig. 27.

1881 Fungia sellata: Quenstedt , p. 871, pl. 177, fig. 25.

1881 Fungia sororiae: Quenstedt , p. 874, pl. 177, fig. 30.

1897 Cyclolites regularis Leymerie : Nötling, p. 11, pl. 1, figs 13–14.

v1903a Cyclolites placenta Reuss : Felix, p. 190.

v1903 a Cyclolites discoidea Blainville : Felix, p. 190.

v1903 a Cyclolites hemisphaerica Michelin : Felix, p. 192, pl. 18, fig. 2.

v1903a Cyclolites polymorpha (Goldfuss) : Felix, p. 198.

1910 Cyclolites polymorpha, Goldfuss ; sp: Pratz, p. 310.

1913 Cyclolites krumbecki: Di Stefani , p. 264, pl. 23, fig. 6.

1914 Cyclolites placenta Reuss : Felix, pars 7, p. 191.

1914 Cyclolites polymorpha (Goldfuss) : Felix, pars 7, p. 191.

1914 Cyclolites regularis Leymerie : Felix, pars 7, p. 192.

v1930 a Cyclolites placenta Reuss : Oppenheim, p. 89 (topotypes studied).

v1930 a Cyclolites weissermeli n. sp.: Oppenheim, p. 99, pl. 5, fig. 11 (topotypes studied).

v1930 a Cyclolites michelini nom. mut.: Oppenheim, p. 100, pl. 40, fig. 2, pl. 46, fig. 13 (topotypes studied). v1930 a Cyclolites pulchellus n. sp.: Oppenheim, p. 105, pl. 51, Fig. 8, pl. 39, figs 9–10a (topotypes studied). v1930 a Cyclolites undulatus var. plana: Oppenheim , p. 106, pl. 12, figs 2–2a (topotypes studied).

v1930 a Cyclolites sororiae (Quenstedt) : Oppenheim, p. 121, pl. 3, fig. 5 (topotypes studied).

1939a Cyclolites senessei n. sp.: Alloiteau, p. 11, pl. 1, figs 12, 12a.

1941 Cyclolites reussi de Fromentel : Alloiteau, p. 71, pl. 30, figs 3–4.

1942 Cyclolites krumbecki De Stefani : Marini, p. 78, pl. 4, figs 1–3, pl. 5, figs 1–12.

1942 Cyclolites krumbecki var. orfellensis: Marini , p. 80, pl. 5, figs 13–18.

?1945 Cyclolites placenta Reuss 1854 : Bataller, p. 39.

?1945 Cyclolites polymorpha Goldfuss sp. 1826: Bataller, p. 39.

?1945 Cyclolites regularis Leymerie 1881 : Bataller, p. 40.

?1945 Cyclolites Reussi Fromentel 1863 : Bataller, p. 40.

1957 Plesiocunnolites goldfussi: Alloiteau , p. 349.

1957 Cunnolites plani-alpici: Alloiteau , p. 349, pl. 9, fig. 4, pl. 11, figs 4, 5, 9.

1957 Cunnolites meringonensis: Alloiteau , pl. 3, figs 9–10, 16–17.

1964 Plesiocunnolites dispar (Quenstedt) : Beauvais, p. 535, pl. 15, figs 1a–c.

1964 Plesiocunnolites sellatus (Quenstedt) : Beauvais, p. 537, pl. 15, figs 4a–d.

1964 Cunnolites sororius (Quenstedt) : Beauvais, p. 538, pl. 16, figs 2a–d.

1964 Plesiocunnolites reussi (Fromentel) : Beauvais, p. 543, pl. 16, figs 5a–c.

1970 Cunnolites plani-alpici Alloiteau : Tschechmedjeva, p. 37, pl. 1, figs 3–4.

1978 Cunnolites (Plesiocunnolites) dispar (Quenstedt) : Turnšek, p. 121, pl. 27, figs 1–4.

1981 Cyclolites regularis Leymerie : Abed & El-Asaad, p. 285, pl. 2, figs 3a–b.

v1982 Cunnolites placenta (Reuss) 1854 : Beauvais, vol. 2, p. 163, pl. 40, fig. 7, pl. 41, fig. 1.

v1982 Cunnolites senessei (Alloiteau) 1939 : Beauvais, vol. 2, p. 165 (topotypes studied).

v1982 Cunnolites sellatus (Quenstedt) 1881 : Beauvais, vol. 2, p. 169 (topotypes studied).

v1982 Cunnolites sellatus nefgrabenensis nov. s. sp.: Beauvais, vol. 2, p. 171, pl. 40, fig. 8 (topotypes studied).

v1982 Cunnolites reussi (de Fromentel) 1870 : Beauvais, vol. 2, p. 184, pl. 41, fig. 6 (topotypes studied).

v1982 Cunnolites plani-alpici Alloiteau 1957 : Beauvais, vol. 2, p. 189, pl. 41, figs 7–8 (topotypes studied).

v1982 Cunnolites undulatus plana (Oppenheim) 1930 : Beauvais, vol. 2, p. 198.

v1982 Cunnolites undulatus (Goldfuss) 1826 : Beauvais, vol. 2, p. 198 (older synonyms cited therein) (topotypes studied).

v1982 Paracunnolites (?) discoideus (Goldfuss) 1826: Beauvais, vol. 2, p. 207, pl. 42, fig. 6 (topotypes studied).

1986 Cunnolites reussi (de Fromentel, 1870) : Tchechmédjiéva, p. 63ff.

1986 Cunnolites sororius (Quenstedt, 1880) : Tchechmédjiéva, p. 64ff.

1992 Cunnolites polymorphus ( Goldfuss, 1826) : Turnšek, p. 167, fig. 2.

1992 Cunnolites sororius (Quenstedt 1880) : Turnšek, p. 167, fig. 2.

1992 Cunnolites ellipticus (Guettard, 1770) : Turnšek, p. 167, fig. 2.

(v)1997 Cunnolites (Cunnolites) reussi (Fromentel) : Turnšek, p. 53, pl. 53, figs A–H.

(v)1997 Cunnolites (Cunnolites) sellata (Quenstedt) : Turnšek, p. 54, pl. 54, figs A–C.

(v)1997 Cunnolites (Plesiocunnolites) dispar (Quenstedt) 1880 : Turnšek, p. 58, pl. 58, figs A–D.

(v)1997 Cunnolites (Plesiocunnolites) undulata (Goldfuss) 1826 : Turnšek, p. 62, pl. 62, figs A–D.

v1998 Cunnolites sp. : Baron-Szabo, p. 152, pl. 4, Fig. 1, pl. 8, Fig. 4, Tab. 4.

2001 Cunnolites regularis ( Leymerie 1878) : Löser & Liao, p. 666–667.

2002 Cyclolites discoidea ( Goldfuss 1826) : Löser et al., p. 189 (older synonyms cited therein).

2002 Cyclolites polymorpha ( Goldfuss 1826) : Löser et al., p. 204 (older synonyms cited therein).

2002 Cyclolites regularis Leymerie 1878 : Löser e t al., p. 206 (older synonyms cited therein).

2002 Cyclolites reussi (de Fromentel 1863) : Löser et al., p. 207 (older synonyms cited therein).

non 2002 Cyclolites undulata undulata ( Goldfuss 1826) : Löser et al., p. 214 (older synonyms cited therein).

2002 Cunnolites dispar ( Quenstedt, 1881) : Baron-Szabo, p. 143.

2002 Cunnolites ellipticus (Guettard, 1770) : Baron-Szabo, p. 143.

2002 Cunnolites plani-alpici Alloiteau 1957 : Baron-Szabo, p. 143.

2002 Cunnolites michelini (Oppenheim, 1930) : Baron-Szabo, p. 143.

2002 Cunnolites meringonensis Alloiteau 1957 : Baron-Szabo, p. 143.

2002 Cunnolites placenta ( Reuss, 1854) : Baron-Szabo, p. 143.

v2002 Cunnolites polymorpha ( Goldfuss, 1826) : Baron-Szabo, p. 143, pl. 102, fig. 6, pl. 103, fig. 2.

2002 Cunnolites pulchellus (Oppenheim, 1930) : Baron-Szabo, p. 143.

2002 Cunnolites regularis ( Leymerie, 1878) : Baron-Szabo, p. 143.

2002 Cunnolites senessei Alloiteau, 1939 : Baron-Szabo, p. 143.

2002 Cunnolites sellatus ( Quenstedt, 1881) : Baron-Szabo, p. 143.

2002 Cunnolites sellatus nefgrabenensis Beauvais, 1982 : Baron-Szabo, p.143.

2002 Cunnolites sororius ( Quenstedt, 1881) : Baron-Szabo, p. 143.

v2002 Cunnolites undulatus ( Goldfuss, 1826) : Baron-Szabo, p. 143, pl. 102, fig. 2.

2002 Cunnolites undulatus plana Beauvais, 1982 : Baron-Szabo, p.143.

2002 Cunnolites weissermeli (Oppenheim, 1930) : Baron-Szabo, p. 143.

v2004 Cunnolites polymorpha (Goldfuss) : Moosleitner, p. 175, pl. 73, figs 3, 4, 6, 12, 13.

Dimensions. d (min)/d (max)=0.7–0.98; s/mm=17–20/ 5 in peripheral areas, up to 25/ 5 in peripheral areas in growing stage, 17–20/5 around calicinal pit. corallum.

Remarks. In having a corallite that varies substantially in outline throughout ontogeny (measurements were taken from type material and illustrations of type material), the specimens examined, as listed in the synonymy, show close affinities to C. giganteus and C. tenuiradiata , but they are distinguished from C. giganteus by having septal densities that are significantly larger, and differ from C. tenuiradiata by having septal densities that are both equal and larger throughout the corallum.

Type locality of species. Cretaceous of France (Dauphiné, Haute Alpes) .

Distribution. Cretaceous of France and Spain, Albian of England (Norfolk, Hunstanton), Lower Cenomanian of Tibet, Upper Cretaceous of Romania, Italy, and Turkey, Lower Senonian of Portugal, Santonian- Campanian of Slovenia, Romania, and Austria (Gosau Group), Campanian of Germany and Hungary, Upper Campanian of Serbia (Fruska Gora) and Bulgaria, Campanian-Maastrichtian of Serbia and Tibet (Yadong), Campanian-Lower Maastrichtian of central Saudi Arabia, Maastrichtian of Austria, Pakistan, and Libya, Paleocene of India.

Cunnolites angiostoma ( Kühn, 1933)

Pl. 16, Fig. 1

v*1933 Cyclolites angiostoma nov. spec.: Kühn, p. 179, pl. 17, figs 5–6.

1942 Cyclolites elliptica Lamarck (Guettard sp.): Maccagno, p. 793, pl. 1, figs 5–5b.

2002 Cyclolites angiostoma, Kühn 1933 : Löser et al., p. 185.

2002 Cunnolites angiostoma ( Kühn, 1933) : Baron-Szabo, p. 143.

Dimensions. d (min)/d (max)=0.85–0.89; d (min)= 25–28 mm; d (max)= 28–33 mm; s/mm=20–25/ 5 in peripheral areas, 25/5 around calicinal pit.

Description. Solitary, free, somewhat elliptical in outline; septa thin, equal, densely packed; septal density nearly equal throughout the corallum but slightly larger around the calicinal pit; calicinal pit elongate.

Type localit y of species. Maastrichtian of Iran ( Neyriz , Esfahan) .

Distribution. Maastrichtian of Iran and Somalia.