Deltochilum (Calhyboma) arrowi Paulian, 1939

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 81-83

Deltochilum (Calhyboma) arrowi Paulian, 1939 stat. n. Plates 19C, 57A

Deltochilum (E.) tessellatum var. Arrowi Paulian, 1939: 18 (original description. Type locality: Équateur [= Ecuador], Río Pescado).

Deltochilum tessellatum var. arrowi : Blackwelder 1944: 203 (list of species from Latin America); Pereira and D’Andretta 1955a: 43 (redescription); Chamorro et al. 2018: 94 (cited for Ecuador).

Deltochilum var. arrowi : Contreras 1951: 222 (cited for Colombia).

Types specimens.

Deltochilum (E.) tessellatum var. Arrowi Paulian, 1939. The holotype (♂) and one paratype are deposited at the NHML (see Paulian 1939: 18). Locality: Ecuador Río Pescado. Examined.

Holotype (♂): "Ecuador / Río- / Pescado [p]", "v-17-1922 / 15000 ft / GHTate [p]", "Frank R. Manson / Collection [p]", "Ex coll. / F.R.Manson. / Brit. Mus. / 1923- 141 [p]", "Deltochilum / tessellatum / var. arrowi Paulian / Type [hw]", "Holo- / type [p, red margin]".

Paratype (♀): "Ecuador / Río- / Pescado [p]", "v-17-1922 / 15000 ft / GHTate [p]", "Ex coll. / F.R.Manson. / Brit. Mus. / 1923-141 [p]", "Para- Type [p, yellow margin]".

Distribution.

Colombia and Ecuador.

Records examined.

CAÑAR: Javin, 1400 m (4 specimens CEMT). COTOPAXI: San Francisco de las Pampas (1 specimen CEMT). MANABÍ: Río Pescado (2 specimens NHML). PICHINCHA: E B La Hesperia, 1200 m (1 specimen CEMT); Pacto, 1000 m (1 specimen CEMT).

Temporal data.

Collected in January, February, March, and October.

Remarks.

Inhabits coastal evergreen foothill forests from 850-1200 m a.s.l. Collected using pitfall traps baited with pig feces.

Paulian in 1939 described Deltochilum (E.) tessellatum var. arrowi (written Arrowi , type locality Río Pescado, Ecuador) as a variety of Deltochilum tessellatum Bates, 1870 (type locality Gualaquiza, Ecuador, Amazonian region), which made the former name available as a subspecific category (see: ICZN 1999, Article 45.6.4). However, upon examining the external and genital morphology of the type specimens of D. tessellatum arrowi (holotype ♂, deposited at the NHML, Plate 57A) and D. tessellatum (lectotype ♂ here designated, deposited at the MNHN, Plate 57B) (specifically, the differences observed in the elytral striae and shape of the aedeagus) we determined them to be two distinct species. Therefore, to maintain the subspecific name originally proposed by Paulian (1939), we elevate the variation arrowi to species level as follows: Deltochilum (Calhyboma) arrowi Paulian, 1939 stat. n. The holotype is here illustrated (♂ Plate 57A) and originates from the locality Río Pescado, Ecuador ( Manabí province, in the coastal region).