Diplommatina cyclostoma Möllendorff, 1897

Hausdorf, Bernhard, 2017, Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java, Zootaxa 4312 (2), pp. 201-245 : 212

Diplommatina cyclostoma Möllendorff, 1897

( Figs 13 , 25 ; Table 1)

Diplommatina (Sinica) cyclostoma Möllendorff, 1897: 95 (JaVa) . Kobelt 1902: 462. Zilch 1953: 28, pl. 12 fig. 177.

Diplommatina cyclostoma — Van Benthem Jutting 1948: 590, fig. 47.

Diplommatina auriculata —Heryanto 2008: 362 (partly, non Möllendorff, 1897). Heryanto 2009: 332 (partly, non Möllendorff, 1897).

Diplommatina sulcicollis —Heryanto 2008: 362 (partly, not Möllendorff 1897). Heryanto 2009: 332 (partly, not Möllendorff 1897).

Palaina gedeana —Heryanto 2008: 362 (partly, non Möllendorff, 1897). Heryanto 2009: 332 (partly, non Möllendorff, 1897).

Diagnosis. Diplommatina cyclostoma is characterized by the spindle shaped, very closely ribbed shell with a circular peristome.

Shell ( Fig. 25 ; Table 1). Dextral; spindle-shaped with conical early whorls and a narrowed last whorl; with 5.5–6 whorls separated by a deep suture; protoconch smooth; teleoconch with fine, oblique ribs (84–116 on the penultimate whorl; fewer on the last whorl); without discernible spiral striae between the ribs; light corneous; subtranslucent; last whorl rounded, with an indistinct ventral constriction; aperture circular; upper insertion of peristome distinctly ascending towards aperture; insertions of the peristome connected by a distinct callus; the peristome slightly expanded, thickened, not doubled or only doubled on columellar and basal side; aperture with a low columellar lamella, which is sometimes missing; without parietal lamella; with a short spiral palatal fold ventrolaterally left inside last whorl in the middle of the whorl and with a long radial palatal fold ventrally; umbilicus closed; operculum thin, corneous, circular, paucispiral. Shell height 2.9–3.25 mm, shell diameter 1.6– 1.95 mm.

Remarks. Diplommatina cyclostoma differs from D. baliana , which was erroneously synonymized with D. cyclostoma by Vermeulen & Whitten (1998), in the larger shell with close-set ribs without discernible spiral striae between the ribs.

Habitat. In leaf litter and on the vegetation in humid forests at 2000–3000 m a.s.l.

Type material. Indonesia, Java, West Java: Cikuray (Tjikorai), 7°03'S 107°20'E (lectotype of D. cyclostoma [designated by Zilch 1953] SMF 105214; paralectotypes of D. cyclostoma SMF 105215/16).

Other material. Indonesia, Java (SMF 192583/4).—West Java: Cianjur, Kandang Badak, saddle between Mt. Pangrango and Mt. Gede, 2400 m a.s.l., 6°47'S 106°58'E (ZMA 68411/100, vouchers to van Benthem Jutting 1948); Cianjur, Mt. Pangrango near Cibodas, 2400 m a.s.l., 6°46'S 106°58'E (ZMA 68413/27, 68415/77, vouchers to van Benthem Jutting 1948); Cianjur, slope of Mt. Pangrango, 2900 m a.s.l., 6°46'S 106°58'E (ZMA 68412/16); Cianjur, Mt. Pangrango, 3000 m a.s.l., 6°46'S 106°58'E (SMF 105217/7, vouchers to Zilch 1953); Cianjur, Mt. Gede, 2500 m a.s.l., 6°47'S 106°58'E (van Benthem Jutting 1948); Cianjur, Mt. Gede, along the road between Cibeurem and Kandang Badak, 2000 m a.s.l., 6°46'S 106°59'E (ZMA 68414/1, vouchers to van Benthem Jutting 1948); Cianjur, Mt. Gede, 2300–2500 m a.s.l., 6°46'S 106°59'E (SMF 105218/9, vouchers to Zilch 1953); Garut, Mt. Cikuray, 2500 m a.s.l., 7°19'S 107°51'E (SMF 105216/1); Garut, Mt. Cikuray, 2500 m a.s.l., 7°19'S 107°51'E (SMF 105216/1); Garut, Mt. Cikuray, 2600 m a.s.l., 7°19'S 107°52'E (van Benthem Jutting 1948); Majalengka, Argamukti, Mt. Ciremai National Park, 6°54'S 108°22'E (MZB 17976/1, voucher to Heryanto 2008, 2009 [as D. sulcicollis ]; MZB 17977/3, vouchers to Heryanto 2008, 2009 [as D. auriculata ]); Majalengka, Argamukti, Mt. Ciremai National Park, 1735 m a.s.l., 6°54'S 108°22'E (MZB 17973/3, vouchers to Heryanto 2008 [as P. gedeana ]).—Central Java: Dieng Highland, Mt. Prahu, 2500 m a.s.l., 7°11'S 109°55'E (ZMA 68410/2, vouchers to van Benthem Jutting 1948).

Distribution ( Fig. 13 ). Diplommatina cyclostoma is endemic to West and Central Java. Records from Madura and Bali ( Vermeulen & Whitten 1998) refer to D. baliana . A record from Sumatra (Heryanto 2013) is also questionable.

Figures

FIGURES 10 – 14. DiStribution of Diplommatinidae in JaVa. 10. Diplommatina planicollis Möllendorff, 1897, Diplommatina heteroglypha Van Benthem Jutting, 1948 and Diplommatina javana Möllendorff, 1897. 11. Diplommatina halimunensis n. sp., Diplommatina kakenca n. sp. and Diplommatina heryantoi n. sp. 12. Diplommatina perpusilla Möllendorff, 1897 and Diplommatina ristiae n. sp. 13. Diplommatina sulcicollis Möllendorff, 1897 and Diplommatina cyclostoma Möllendorff, 1897. 14. Diplommatina calcarata Möllendorff, 1897 and Diplommatina tetragonostoma Möllendorff, 1897.

FIGURES 23 – 25. Diplommatina Spp. from JaVa. 23. Diplommatina heteroglypha Van Benthem Jutting, 1948, EaSt JaVa: Banyuwangi, Mt. Ijen, Merapi Ungup-ungup (paratype ZMA 135947). 24. Diplommatina javana Möllendorff, 1897, JaVa (lectotype SMF 105202). 25. Diplommatina cyclostoma Möllendorff, 1897, WeSt JaVa: Cikuray (lectotype SMF 105214). Scale bar = 1.0 mm.