Gayennoides molles, RAMÍREZ, 2003

RAMÍREZ, MARTÍN J., 2003, The Spider Subfamily Amaurobioidinae (Araneae, Anyphaenidae): A Phylogenetic Revision At The Generic Level, Bulletin of the American Museum of Natural History 2003 (277), pp. 1-262 : 125-126

Gayennoides molles , new species Figures 64B , 65

TYPES: Female holotype and male paratype from Chile, Región V, Petorca province, Los Molles, 2 m, under succulent rock cover along coast, ca. 32°14̍S, 71°30̍W, 9.I.1985, N. Platnick, O. Francke , deposited in AMNH.

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Similar to G. losvilos , but distinguished by having only two teeth on the cheliceral retromargin, and a larger, rectangular epigynal median field.

FEMALE (holotype): Total length 10.65. Carapace length 4.05, width 2.93, wider on legs II–III. Length of tibia/metatarsus: I, 2.80/2.40; II, 2.63/2.33; III, 2.10/2.27; IV, 2.97/3.46. Palpal tarsus length 1.25. Chelicerae unmodified, with two teeth on retromargin. Sternum length 2.10, width 1.60. Spines: leg I, femur d 1–1–1, p 2ap; tibia v 2–2–2; metatarsus v 2bas. II, femur d 1–1–1, p 0­ d1­ d1; tibia v r1–2–2, p 0–1; metatarsus v 2bas. III, femur d 1–1–1, p and r 0­ d1­ d1; patella r 1; tibia v p1­ p1–2, p 1­ d1­ 1­ 0, r d1– 1, d r1bas; metatarsus v 2–2–2, p and r d1– 1–1, d 0­ p1–2. IV, femur d 1–1–1, p 0­ d1­ d1, r d1ap; patella r 1; tibia v p1–2–2, p and r 1­ d1­ 1­ 0, d r1bas; metatarsus = III. Abdomen length 5.72, width 3.05, spiracle–epigastrium 2.57, spiracle–spinnerets 0.77. Col­ or (fig. 65A): carapace reddish brown, with lateral dark stripes. Chelicerae reddish brown. Legs pale grayish, with very dark dots at dorsal and lateral spine bases, some spots on patellae and tibiae. Sternum grayish, with tenuous dark spots in front of coxae I– III, median dark spot. Abdomen yellow with dark spots, anterior half of cardiac area gray, two large spots at insertions of posterior dorsoventral muscles, several median spots extending extending to anal tubercle. Venter pale with a few dark dots. Epigyne (fig. 65F, G): median field rectangular. Copulatory ducts joining at posterior margin.

MALE (paratype): Total length 7.60. Carapace length 3.75, width 2.80. Length of tibia/ metatarsus: I, 3.85/3.35; II, 3.13/2.90; III, 2.30/2.50; IV, 3.17/3.53. Sternum length 2.00, width 1.57. Chelicerae longer than those of female, median promarginal tooth quite larger, closer to apical tooth. Spines as in female, except: leg II, metatarsus p 1–0. III, tibia v p1–2–2 or p1­ p1–2. Abdomen length 4.40, width 2.00, spiracle–epigastrium 1.57, spiracle–spinnerets 0.60. Color as in female (fig. 64B). Palp (fig. 65B–E): tibia long, width/length 0.55, cymbium with conspicuous notch at retrolateral basal margin. Copulatory bulb: paramedian apophysis with blunt cusp, two ridges at sides.

NATURAL HISTORY: Many specimens collected under the Crassulaceae lining stones near seashore.

DISTRIBUTION: Chilean coast, from Antofagasta to Quillota provinces. Also collected in two noncoastal localities, Cuesta La Dormida and Cuesta El Melón, in Region V. This apparently disjunct distribution is similar to that of Acanthoceto ladormida .

OTHER MATERIAL EXAMINED: CHILE: Región II (Antofagasta): Antofagasta: Caleta Hueso, W of Taltal, 28.I.1941, Junius Bird, 1♀ 1 immature (AMNH); 4 km N Paposo , 20–50 m, 11.X.1992, N. Platnick, P. Goloboff, K. Catley , 9♀ 2 immatures (AMNH); Cerro Moreno, 900 m, 18.IV.1992, H. Larrain, 1♀ (AMNH); 6 km E Paposo , 480 m, 12.X.1992, N. Platnick, P. Goloboff, K. Catley , 23 13♀ (AMNH), 520 m, 25°01̍S, 70°27̍W, N. Platnick, K. Catley, R. Calderón, R.T. Allen , 1♀ (AMNH); Paposo, 54 km N Taltal, II.1957, W. Toodt, 1♀ (MHNS 532); Cerro Moreno, 900 m, 8.IV.1992, H. Larrain, 1♀ (AMNH); Taltal, La Quinta (bajo piedras), 5–7.X.1983, M. Elgueta, 1♀ (MHNS 766). Región III (Atacama): Chañaral: N Chañaral, 10.X.1992, L. Peña, 23 1 immature (AMNH); Isla de Chañaral, ca. 4 km from coast, 30.X.1980, L. Peña, 13 (AMNH); Pan de Azúcar, 10–12.X.1992, L. Peña, 1♀ (AMNH), 12.X.1992, L. Peña, 23 1♀ 1 immature (AMNH). Copiapo´: Copiapo´, 6.V.1964, G. Mann F., 1♀ (MHNS); Puerto Viejo, 15–16.X.1992, L. Peña, 13 2♀ 4 immatures (AMNH), 17.IV.1980, L. Peña, 1♀ (AMNH); Puerto Viejo, S La Caldera, 15–16.X.1992, L. Peña, 33 1♀ 2 immatures (AMNH); E of Puerto Viejo, 9–10.X.1980, L. Peña, 1♀ 2 immatures (AMNH). Región IV (Coquimbo): Elqui: Choros Bajos, 21.X.1992, L. Peña, 1♀ (AMNH); 6 km S Cruz Grande, 20 m, 7.X.1992, N. Platnick, P. Goloboff, K. Catley , 7♀ (AMNH); 6.X.1992, N. Platnick, P. Goloboff, K. Catley , 5♀ (AMNH); 9 km S Cruz Grande, beach, 5 m, 11.XI.1993, 29°29̍S, 71°19̍W, N. Platnick, K. Catley, M. Ramírez, T. Allen , 13 1 immature (AMNH), 13 1 immature (MACN­Ar); 16 km S Cruz Grande, 140 m, 7.X.1992, N. Platnick, P. Goloboff, K. Catley , 3♀ (AMNH); Puerto Pichidangui, Isla de los Locos, 19.VII.1961, R. Donoso, A.F. Archer, 1♀ (AMNH); La Serena, 150 m, coastal scrub matorral, 4.XI.1981, N. Platnick, T. Schuh , 1♀ (AMNH); 19 km N La Serena, 150 m, coastal scrub matorral, 1.XI.1980, N. Platnick, T. Schuh , 1♀ (AMNH). Limarı´: Fray Jorge Natl. Park, 400 m, 3.X.1992, N. Platnick, P. Goloboff, K. Catley , 1♀ (AMNH); Termas de Socos, 300 m, 2.X.1992, N. Platnick, P. Goloboff, K. Catley , 1♀ (AMNH). Choapa: Huentelauquén, coastal town, 27.XII.1980, L. Peña, 13 1♀ (AMNH), 3.X.1990, L. Peña, 1♀ (AMNH); 20 km N La Serena (Rt. 5 km 491), 120 m, 7.X.1992, N. Platnick, P. Goloboff, K. Catley , 23 4♀ (AMNH); Los Vilos, 25.IX.1966, E.I. Schlinger, 13 (CAS); 5 km N Los Vilos, 3 m, under succulent rock cover along cove, 5.I.1985, N. Platnick, O. Francke , 3♀ 5 immatures (AMNH); 19 km N Los Vilos, Rt. 5, km 244, 5 m, 9.XI.1993, 31°45̍S, 71°31̍W, N. Platnick, K. Catley, M. Ramírez, T. Allen , 13 1♀ (AMNH); Isla de Los Locos, Pichidangui, 29.IV.1961, R. Donoso, 1♀ 1 immature (AMNH). Coquimbo: Caleta Oscuro, 31°26̍S, 71°37̍W, 23.IX.1966, E.I. Schlinger, 13 8♀ 1 immature (CAS), 2.X.1983, E. Maury, 3♀ (MACN­Ar); 5 km S Coquimbo, 30 m, coastal scrub matorral, 31.X.1981, N. Platnick, T. Schuh , 13 1 immature (AMNH); Corral de Julio, Quebrada La Madera, 3.X.1972, N. Figueroa, 1♀ (UC); La Herradura, 80 m, 3.X.1992, N. Platnick, P. Goloboff, K. Catley , 1♀ (AMNH). Región V (Valparaíso): Petorca: Los Molles, 3.VII.1975, R. Pérez, 13 (UC); Los Molles, Rt. 5, km 188, elev. 10 m, 9.XI.1993, 32°14̍S, 71°30̍W, N. Platnick, K. Catley, M. Ramírez, T. Allen , 13 2♀ (AMNH), 23 1♀ 1 immature (MACN­Ar), 13 1♀ (MHNS, photos MJR 1331–1332), 2 m, under succulent rock cover along coast, 9.I.1985, N. Platnick, O. Francke , 5♀ 10 immatures (AMNH); 13 km S Los Molles, 32°12̍S, 71°27̍W, beach dunes, 22.IX.1966, E.I. Schlinger, 4♀ 2 immatures, 1♀ (CAS); Quebrada del Chivato, 1 km S Los Molles, 10 m, 2.X.1992, N. Platnick, P. Goloboff, K. Catley , 5♀ 3 immatures (AMNH), 30.X.1988, P. Goloboff, E. Maury, C. Szumik, 1♀ 2 immatures (MACN­Ar); Quebrada Huaquén, Pichicuy, 10 m, 2.X.1992, N. Platnick, P. Goloboff, K, Catley , 13 4♀ (AMNH); Cuesta El Melón, 430 m, 8.XI.1993, 32°37̍S, 71°14̍W, N. Platnick, K. Catley, M. Ramírez, T. Allen , 13 (AMNH). Quillota: Cuesta La Dormida (east side), 33°04̍S, 71°02̍W, 750– 1000 m, 20.IX.1966, E.I. Schlinger, 13 2♀ (CAS). No Locality: Loc. 280 (presumably from Chile), L. Peña, 1♀ (IRSN IG 19736).

Figures

Fig. 64. Gayennoides spp. A. G. losvilos, sp. n., female (Elqui, Cruz Grande, photo MJR 1330). B. G. molles, sp. n., male (Petorca, Los Molles, photo MJR 1331).

Fig. 65. Gayennoides molles, n. sp., female holotype, male paratype. A. Male. B. Male copulatory bulb, ventral view. C. Same, retrolateral view. D. Same, apical view. E. Male palp, retrolateral view. F. Cleared epigyne, ventral view. G. Epigyne, ventral view. Scale bars = A, 1 mm; B–G, 0.2 mm.