Granulatocincta callim, Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 48-49

Granulatocincta callim nov. sp.

Figs 17A 1 –A 3 , B 1 –B 3 , C 1 –C 3 , D 1 –D 2 , 3Y , 5 , 6

Pleurotoma rustica Brocc. — Hörnes 1848: 20 [non Granulatocincta rustica ( Brocchi, 1814) ].

Pleurotoma Sotterii Micht. — Hörnes 1854: 338, pl. 36, figs 16a–c [non Granulatocincta sotterii ( Michelotti, 1847) ].

Clavatula sotteri (Michelotti) — Papp 1954: 57, pl. 9, figs 6–7 [non Granulatocincta sotterii ( Michelotti, 1847) ].

Clavatula sotteri Micht. — Jekelius 1944: 86, pl. 23, figs 10–11[non Granulatocincta sotterii ( Michelotti, 1847) ] [non fig. 12 =?].

Clavatula (Clavatula) sotteri (Michelotti) — Švagrovský 1958: 16, pl. 4, fig. 8 [non Granulatocincta sotterii ( Michelotti, 1847) ].

C [lavatula]. (C [lavatula]) sotterii (Micht.) — Sieber 1958a: 157 [non Granulatocincta sotterii ( Michelotti, 1847) ].

Clavatula sotteri (Michelotti) — Boda 1959: 627, pl. 29, fig. 12 [non Granulatocincta sotterii ( Michelotti, 1847) ].

? Clavatula (Clavatula) interrupta apolloniae (Hoernes et Auinger) — Hinculov 1968: 147, pl. 36, figs 12a–b [non ‘Clavatula’ apolloniae ( Hoernes & Auinger, 1891) ].

Clavatula sotteri ( Michelotti, 1847) — Švagrovský 1971: 405, pl. 70, fig. 4 [non Granulatocincta sotterii ( Michelotti, 1847) ].

Clavatula sotteri (Michelotti) — Papp 1974: 354, pl. 10, figs 10–11 [non Granulatocincta sotterii ( Michelotti, 1847) ].

Type material. Holotype: NHMW 1866 /0001/0549, SL: 16.5 mm, MD: 7.9 mm, Wiesen ( Austria), figs 17A 1 –A 3 . Paratype: NHMW 1846 /0037/0351a, SL: 17.6 mm , MD: 8.6 mm, Wiesen ( Austria) , illustrated in Hörnes (1854, pl. 36, figs 16a–c), figs 17C 1 –C 3; NHMW 1846 /0037/0351b, SL: 19.9 mm , MD: 8.9 mm, Wiesen ( Austria) , figs 17B 1 –B 3; NHMW 2021 /0128/0001, SL: 21.5 mm , MD: 9.6 mm, Bad Sauerbrunn ( Austria) , figs 17D 1 –D 2, 3Y. SMF 351868 /5, Bad Sauerbrunn ( Austria) .

Additional studied material. NHMW 2021 /0129/0001, 8 spec., Wiesen ( Austria) .

Type locality. Wiesen ( Austria), Eisenstadt-Sopron Basin .

Type stratum. Mollusc rich sand of the Skalica Formation.

Age. Middle Miocene, Sarmatian ( Ervilia Zone ) (= late Serravallian).

Diagnosis. Granulatocincta species of small size, broad buccinoid profile, coarsely sculptured, early whorls tripartite, bearing two swollen spiral cords with broad beads at adapical and abapical suture and narrower beaded central cord. Last whorl bearing strong rounded tuberculose cords at mid-whorl and narrower cords on base. Short, swollen twisted fasciole. Short siphonal canal.

Etymology. Callim, Latin for unknown to, concealed from. Referring to its overlooked status.

Description. Shell small, solid, broad buccinoid with conical spire; apical angle 41–48°. Protoconch not preserved. Teleoconch of up to eight flat-sided whorls bearing weakly opisthocline axial ribs with tripartite sculpture. Subsutural cord weakly tubercular, central cord with small beads, suprasutural cord with larger, axially elongated, opisthocline beads. On penultimate whorl a further spiral cord appears just above the suture, mostly covered by last whorl. Suture narrowly impressed, linear. Last whorl 65% of total height, ovoid. Subsutural collar broad, sharply delimited, subobsoletely tuberculose. Subsutural ramp narrow, filled with narrow beaded cord. Shoulder delimited by broad cord bearing low tubercles, two slightly narrower broad bands below with low tubercles, lower representing peribasal band. Base weakly constricted, bearing 4–5 narrower beaded cords; all cords separated by narrow grooves and secondary cords may be intercalated in some interspaces. Siphonal fasciole short, swollen, strongly twisted. Aperture wide, ovate; outer lip not thickened by varix, smooth within. Anal sinus deep, wide asymmetrically U-shaped, apex coinciding with beaded cord mid-ramp. Siphonal canal short, broad, shallowly notched at tip. Columella weakly excavated in upper third, weakly twisted below, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming broad callus rim.

Discussion. This species was misidentified as Granulatocincta sotterii ( Michelotti, 1847) by all authors since Hörnes (1854), except for Landau et al. (2013: 287), who doubted this identification. Granulatocincta sotterii was described from the Tortonian of Italy ( Bellardi 1877; Montanaro 1937; Ruggieri & Davoli 1984, Davoli 1990) and is clearly similar to the Paratethyan species in its general appearance. Nevertheless, the Italian species differs in its more conical spire, its constricted base, longer and narrower siphonal canal and the absence of a fourth row of beads at the abapical suture on the penultimate whorl (see Montanaro 1937: 117, pl. 5, figs 2–18; Ruggieri & Davoli 1984, pl. 4, figs 1, 6, 13, 14, 18, 19, Davoli 1990, pl. 8, figs 2–8). If both species are directly related, Granulatocincta sotterii would be the rare case of a Proto-Mediterranean species, which originated in the endemic fauna of the Sarmatian Paratethys. The ancestor of G. callim might be found in the Badenian G. contorta Švagrovský, 1958 .

Paleoenvironment. Inner neritic sand bottoms from littoral to shallow sublittoral environments (own data).

Distribution in Central Paratethys. Sarmatian (middle Miocene): Vienna Basin: Strážnice (the Czech Republic) ( Švagrovský 1958, 1971); Eisenstadt-Sopron Basin: Wiesen, Bad Sauerbrunn ( Austria) ( Papp 1954; hoc opus); Cserhát Mountains ( Hungary) ( Boda 1959), Soceni ( Romania) ( Jekelius 1944).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia).

FIGURE 17 A1–A3. Granulatocincta callim nov. sp., holotype, NHMW 1866/0001/0549, Wiesen (Austria). B1–B3. Granulatocincta callim nov.sp.,paralectotype,NHMW 1846/0037/0351b, Wiesen (Austria).C1–C3. Granulatocincta callim nov. sp., paralectotype, NHMW 1846/0037/0351, Wiesen (Austria). D1–D2. Granulatocincta callim nov. sp., paralectotype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). E. Granulatocincta carasiensis (Jekelius, 1944), MNG P– 4965, Soceni (Romania). F1–F2. Granulatocincta contorta (Švagrovský, 1958), holotype, Z 3076, Kuzmice (Slovakia). G1–G2. Granulatocincta contorta (Švagrovský, 1958), paratype, SNMB Z 3074, Kuzmice (Slovakia). H1–H3. Granulatocincta pelliscrocodili nov. sp., holotype, RGM 776 907, Gödet Creek near Tilkikaya (Turkey). I1–I2. Granulatocincta pelliscrocodili nov. sp., paratype, RGM 776 906, southwest bank of Gödet Creek near Tilkikaya (Turkey). J1–J3. Granulatocincta nataliae (Hoernes & Auinger, 1891), holotype, NHMW 1855/0035/0021, Belgrade/Rakovica (Serbia).