Hadrurinae Stahnke, 1974

Soleglad, Michael E. & Fet, Victor, 2003, High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni), Euscorpius 2003 (11), pp. 1-175 : 111

Subfamily Hadrurinae Stahnke, 1974

Type Genus. Hadrurus Thorell, 1876 .

Composition. This monotypic subfamily includes a

single genus Hadrurus .

Distribution. North America ( USA and Mexico).

Taxonomic history. The content of Hadrurinae is changed here since we transfer the genus Anuroctonus to Chactidae (subfamily Uroctoninae ), so that the subfamily is now monotypic.

Biogeographic history. Species of Hadrurus are the largest desert scorpions in North America, and their evolution clearly is connected to aridization of this continent (Morafka et al., 1994). DNA phylogeny of this genus ( Fet et al., 2001) agrees with its Mexican origin as suggested earlier ( Soleglad, 1976a).

Diagnosis. Synapomorphies. Major neobothriotaxy present on ventral aspect of pedipalpal chela, and on ventral and external aspects of patella; ventral surface of leg tarsus with fused spinule cluster groups; leg tarsus unguicular spur well-developed; genital papillae of male absent. Important Symplesiomorphies. Chelal trichobothrium Et 5 positioned on fixed finger; dorsal edge of cheliceral movable finger with two subdistal denticles; leg coxae IV elongated; lateral carinae partially present on metasomal segment IV; chela with “10-carinae” configuration.

Discussion. See discussion above under subfamily Uroctoninae for the reasons for removing Anuroctonus from Iuroidea .