Phyllobaenus cyanipennis (Gorham, 1883: 175)

Leavengood, John M. & Garner, Beulah H., 2014, Nomenclatural notes on some checkered beetle (Coleoptera: Cleridae) types of the Natural History Museum, London (BMNH), Zootaxa 3760 (3), pp. 301-335 : 306-307

Phyllobaenus cyanipennis (Gorham, 1883: 175)

( Fig. 7 )

Original combination. Hydnocera cyanipennis

Types. Gorham identified the complete type series to comprise eleven specimens, nine of which are from the type locality. Four from Calderas and the one specimen from the Quiche Mountains are in the BMNH, leaving five from Calderas and the one specimen from Capetillo in one or more other museums. Döbler (1982: 416) recognized two of the Calderas syntypes in the SDEI.

Series locality. Calderas, Quiche Mountains and Capetillo, Guatemala.

Type locality. Calderas, Guatemala.

LECTOTYPE / PARALECTOTYPE (here designated): Type [round, red-bordered label], Calderas, Guatemala, Champion, Type, B. C. A. Coll. III (2) Hydnocera cyanipennis Gorham , Hydnocera cyanipennis Gorham ( BMNH; 2 sharing a card mount, “LT” is written below the lectotype, the other specimen is a paralectotype). PARALECTOTYPES: Calderas, Guatemala, Champion, B. C. A. Coll. III (2) Hydnocera cyanipennis Gorham ( BMNH; 2 sharing a card mount); Quiche Mts., 7 – 9000 ft., Champion, B. C. A. Coll. III (2) Hydnocera cyanipennis Gorham , Hydnocera cyanipennis Gorham ( BMNH; 1).

Phyllobaenus cylindricollis (Gorham, 1886: 343) ( Fig. 8 )

Original combination. Hydnocera cylindricollis

Types. Gorham vaguely referenced “about a dozen examples,” six of which are in the BMNH and one in the MNHN. When Gorham (1883) discussed Hydnocera bituberculata Chevrolat he had included both of these species. Two specimens originally considered to be bituberculata Chevrolat by Gorham (1883) were included in the type series for cylindricollis . There are several remaining unrecognized syntypes in other museums. Döbler (1982: 416) considered two SDEI specimens from Bugaba (collected by Champion; not necessarily making it part of the type series) and Teapa, Tabasco (collected by H. H. Smith in January) to be syntypes, the latter of which is most confusing considering Teapa was not included in Gorham’s locality list. Until they can be examined, these two specimens are here excluded from the type series.

Series locality. Chontales, Nicaragua, and Bugaba, David and Volcan de Chiriqui, Panama.

Type locality. Bugaba, Panama.

LECTOTYPE (here designated): Type [round, red-bordered label], Bugaba, Panama, Champion, Sp. figured, Hydnocera cylindricollis, B. C. A. Col. III (2), Hydnocera cylindricollis Gorham ( BMNH; 1). PARALECTOTYPES: Chontales, Nicaragua, Janson, B. C. A. Col. III (2), Hydnocera cylindricollis Gorham ( BMNH; 1); David, Panama, Champion, B. C. A. Col. III (2), Hydnocera cylindricollis Gorham ( BMNH 1); V. de Chiriqui, 800 – 1500 ft, Champion, B. C. A. Col. III (2), Hydnocera cylindricollis Gorham ( BMNH; 1); Bugaba, Panama, Champion, Sharp Coll. 1905-313, Hydnocera bituberculata ( BMNH; 2); Bugaba, 800 – 1500 ft, Champion, H. cylindricollis , Museum Paris Coll Gorham 1914 [blue label], Museum Paris ex. Coll. R. Oberthur [blue label] ( MNHN; 2, sharing a single cardmount).

Phyllobaenus flavifemoratus ( Gorham, 1877: 261) ( Fig. 11 )

Original combination. Hydnocera flavifemorata

= Phyllobaenus pallipes ( Gorham, 1877: 261) [ Hydnocera pallipes ], new synonymy

= Phyllobaenus rufithorax ( Gorham, 1877: 262) [ Hydnocera rufithorax ], new synonymy

Comments. There are no structural differences in the specimens in these three type series. The characters described to distinguish pallipes ( Fig. 12 ) from flavifemoratus fall within the range of variation observed in flavifemoratus . Regarding color, flavifemoratus varies considerably (as do most hydnocerine species) with the legs commonly distinctly banded or faded and largely pale, and the elytral maculae of varying shape and size. The head, thorax and elytra are broadly black, with or without a bluish tinge, or varyingly mottled testaceous to brownishorange, even reddish-orange as described in rufithorax ( Fig. 13 ), the most extreme example yet examined. The elytral apex may be more rounded, evenly weakly serrate, or slightly truncated (less rounded) with an occasional gap in the serration at the apex, or of intermediate form. All three species ( Figs. 11 – 13 ) were described in the same publication, so as the first revisers we fix the precedence of flavifemoratus , selecting it because it was described earliest in the article.

Types. Others syntypes may exist.

Series/ Type locality. Amazon.

LECTOTYPE (here designated): Type [round, red-bordered label], Type, Gorham Type [red label], Amazon, Bates, Fry Coll. 1905.100., H. flavifemorata (BMNH; 1). PARALECTOTYPES: Type, Gorham Type [red label], Amazon, Bates, Fry Coll. 1905.100. (BMNH; 1); 25161 [ Hydnocera flavifemorata Gorh. Type. Amazon. Pará. Bates.], Type, Amazon, Bates, Fry Coll. 1905.100. (BMNH; 1); Type, Amazon, Bates, Fry Coll. 1905.100. (BMNH; 1); Amazon, Bates, H. flavifemorata , Museum Paris Coll Gorham 1914 [blue label], Museum Paris ex. Coll. R. Oberthur [blue label] (MNHN; 1).

Figures

FIGURES 7 – 10. Fig. 7) Phyllobaenus cyanipennis (Gorham), Lectotype [left] and Paralectotype [right] (BMNH); Fig. 8) P. cylindricollis (Gorham), Lectotype (BMNH); Fig. 9) P. impressus (Gorham), Lectotype [left] and Paralectotype [right] (BMNH); Fig. 10) P. intricatus (Gorham), Lectotype [left] and Paralectotype [right] (BMNH).

FIGURES 11 – 16. Fig. 11) Phyllobaenus flavifemoratus (Gorham), Lectotype (BMNH); Fig. 12) P. pallipes (Gorham), Lectotype (BMNH); Fig. 13) P. rufithorax (Gorham), Lectotype (BMNH); Fig. 14) P. pulchellus (Gorham), Lectotype (BMNH); Fig. 15) P. pulchellus (Gorham), Paralectotype (BMNH); Fig. 16) P. trichrous (Gorham), Lectotype (BMNH).