Megaclavatula pilleri, Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 73-74

Megaclavatula pilleri nov. sp.

Figs 22G 1 –G 3 , H 1 –H 3 , 23E 1 –E 3 , F 1 –F 3, 3S , 5 , 6

Pleurotoma (Clavatula) styriaca Auing. — Hoernes & Auinger 1891: 348 (partim), pl. 47, figs 6–9 only [non Perrona styriaca ( Hilber, 1879)].? Pleurotoma styriaca Auing. — Moisescu 1955b: 262 [non Perrona styriaca ( Hilber, 1879) ]. C [lavatula]. (C [lavatula]) styriaca (Auing.) — Sieber 1958a: 157 [non Perrona styriaca ( Hilber, 1879) ].? Clavatula styriaca (Auing.) — Báldi 1960: 81 [non Perrona styriaca ( Hilber, 1879) ]. Clavatula (Clavatula) styriaca ( Hilber, 1879) — Mikuž 1998: 73, pl. 3, figs 3, fig. 4? [non Perrona styriaca ( Hilber, 1879) ]. Clavatula asperulata ( Lamarck, 1822) s.l. —Harzhauser 2002: 118, pl. 10, fig. 18 [non Megaclavatula asperulata ( Lamarck, 1822)].? Clavatula styriaca ( Hilber, 1879) — Mikuž 2009: 32, pl. 10, fig. 139 [non Perrona styriaca ( Hilber, 1879) ].

Type material. Holotype: NHMW 1949 /0005/0051a, Baden-Sooss ( Austria), SL: 39.4 mm, MD: 13.9 mm, figs 22G 1 –G 3, 3S . Paratypes: NHMW 1949 /0005/0051b, Baden-Sooss ( Austria) , SL: 42.9 mm, MD: 16.0 mm, figs 22H 1 –H 3; NHMW 1949 /0005/0051c, Baden-Sooss ( Austria) , SL: 55.1 mm, MD: 20.7 mm, illustrated in Hoernes & Auinger (1891, pl. 47, fig. 7), figs 23E 1 –E 3; NHMW 1949 /0005/0051d, Baden-Sooss ( Austria) , SL: 48.1 mm, MD: 18.8 mm, illustrated in Hoernes & Auinger (1891, pl. 47, fig. 6), figs 23F 1 –F 3; NHMW 2021 /0132/0002, 2 spec., Baden-Sooss ( Austria) , SMF 351870 /1, Baden-Sooss ( Austria) .

Type locality. Baden-Sooss ( Austria), Vienna Basin .

Type stratum. Silt and clay of the Baden Formation.

Age. Middle Miocene, middle Badenian (late Langhian).

Etymology. In honor of Werner E. Piller (University Graz, Austria), in recognition of his contributions on Miocene stratigraphy.

Diagnosis. Shell, medium-sized, solid, moderately broad fusiform with subsutural collar bearing small sharp tubercles. Shoulder and peribasal cords very weak or subobsolete. Strongly constricted base. Spiral sculpture almost absent.

Description. Shell medium-sized, solid, moderately broad fusiform, with moderately high, slightly gradate spire; apical angle ~39–45°. Protoconch not preserved. Teleoconch of ten whorls. Early teleoconch whorls straight-sided, with tripartite spiral sculpture; weak, beaded subsutural cord, even narrower beaded mid-whorl cord, prominent strongly beaded suprasutural cord. Sculpture changing on fourth to fifth teleoconch whorl; subsutural cord develops into prominent swollen collar bearing wide spaced, low pointed tubercles (12–14 on last whorl) placed a short distance below suture; mid-cord disappears, mid-whorl concave. Suprasutural cord covered by subsequent whorl. Spiral sculpture of delicate spiral threads on early teleoconch whorls, weakening or subobsolete on last whorl. Suture narrowly incised, weakly undulating. Last whorl ~65% of total height, broad subsutural collar bearing pointed tubercles, broad, weakly concave subsutural ramp, rounded to slightly angular shoulder. Base strongly constricted. Spiral sculpture almost absent, except for weak to subobsolete shoulder and peribasal cords and a few weak cords over base and siphonal fasciole. Siphonal fasciole moderately long, poorly delimited, with weak growth increments. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus wide, deep, broadly Vshaped, with apex on most concave portion of ramp below collar. Siphonal canal moderately long, slightly recurved and bent to left, unnotched at tip. Columella strongly excavated in upper third, weakly twisted at fasciole, smooth. Columellar and parietal callus moderately thickened, adherent, forming broad callus rim.

Discussion. Megaclavatula pilleri nov. sp. is slightly reminiscent of M. laevigata ( Eichwald, 1830) , which differs in its larger size and in the presence of prominent peribasal and perifasciolar cords. Megaclavatula pilleri is reminiscent of some Perrona species, but differs from Perrona in its tripartite sculpture on early teleoconch whorls. Hoernes & Auinger (1891) and Sieber (1958a) (inter alia) confused this species with Perrona styriaca ( Hilber, 1879) . Perrona rosaliae ( Hoernes & Auinger, 1891) is larger, has the comma-shaped axial riblets on early whorls typical of the genus, has a higher spire, develops a tuberculate basal spiral cord and a U-shaped anal sinus. Perrona francisci ( Toula, 1901) is a similar species from the Serravallian of Turkey with nearly identical sculpture, but differs from M. pilleri in the much longer siphonal canal and its more slender outline (see Toula 1901: 260, pl. 8, fig. 10.; Landau et al. 2013: 284, pl. 47, figs 5–6). Moreover, Perrona francisci has a completely different early teleoconch with opisthocline ribs (see Landau et al. 2013, pl. 73, figs 8–9).

Paleoenvironment. Unknown. The occurrences in the Baden Formation in the Vienna Basin would suggest middle to outer neritic environments but shallow marine species have frequently been transported into deeper environments at the Baden-Sooss locality (own data). Occurrences in the Korneuburg and Krka basins, would rather indicate inner neritic environments (e.g., Harzhauser et al. 2002).

Distribution in Central Paratethys. Karpatian (early Miocene): Korneuburg Basin: Kleinebersdorf ( Austria) (Harzhauser 2002); Badenian (middle Miocene): Vienna Basin: Baden-Sooss ( Austria) ( Hoernes & Auinger 1891), Krka Basin: Gorenja Stara vas, Ivanji dol ( Slovenia) ( Mikuž 2009);? Făget Basin: Lăpugiu de Sus ( Romania) ( Moisescu 1955b).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia).

FIGURE 22 A1–A3. Megaclavatula laevigata (Eichwald, 1830), NHMW 1846/0037/0308, Baden (Austria). B1–B3. Megaclavatula subsculpta (Schaffer, 1912), lectotype, KM Inv. No. F/1125, Maria Dreieichen (Austria). C1–C2. Megaclavatula nemethi (Kovács & Vicián, 2021), holotype, HNHM, PAL 2021.21.1, Letkés (Hungary). D. Megaclavatula nemethi (Kovács & Vicián, 2021), paratype, NHMW 2021/0002/0003, Letkés (Hungary). E1–E2. Megaclavatula? szontaghyi (Strausz, 1926), holotype, M.116, Bán, Baranya County (Hungary). F. Megaclavatula neudorfensis (Schaffer, 1898), holotype, NHMW 2021/0137/0001, Devínska Nová Ves (Slovakia). G1–G3. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). H1–H3. Megaclavatula pilleri nov. sp., paratype, NHMW 1949/0005/0051b, Baden-Sooss (Austria).

FIGURE 23 A1–A3. Megaclavatula polonica (Pusch, 1837), neotype, NHMW 2021/0138/0001, Korytnica (Poland). B1–B3. Megaclavatula polonica (Pusch, 1837), NHMW 2021/0138/0002, Korytnica (Poland).C1–C3. Megaclavatula polonica (Pusch, 1837),NHMW2021/0139/0001,Baden-Sooss (Austria).D1–D3.Megaclavatula polonica(Pusch,1837),NHMW1862/0001/0324, Bad Vöslau (Austria). E1–E3. Megaclavatula pilleri nov. sp., paratype, NHMW 1949/0005/0051c, Baden-Sooss (Austria). F1–F3. Megaclavatula pilleri nov. sp., paratype, NHMW 1949/0005/0051d, Baden-Sooss (Austria).