Eunatalis porcata (Fabricius, 1787)

Gerstmeier, Roland & Seitner, Martin, 2013, Revision of the checkered beetle genus Eunatalis Schenkling, 1909 (Coleoptera: Cleridae: Clerinae), Zootaxa 3698 (1), pp. 1-77 : 49-52

Eunatalis porcata (Fabricius, 1787)

( Figures 39–40 , 57 C)

Notoxus porcatus Fabricius, 1787: 127 ; Opilus porcata Klug, 1842: 318 ; Natalis porcata Waterhouse, 1894: 286 ; Ekis 1975: 23.

Natalis cribricollis Spinola, 1844: 203 (t. 16, f. 4); Chevrolat 1876: 4; Gorham 1876: 71; Blackburn 1899: 32. Natalis inconspicua Blackburn, 1890: 124 ; Blackburn 1899: 34.

Specimens examined: Notoxus porcatus : Lectotype (designated here): Not. Porcatus , F. Mant. Ins. n. (BMNH).

Natalis cribricollis : Holotype: Holotype, Eunatalis cribricollis (Spinola) ; MRSN—Torino Coll., Spinola Coleoptera box 419 (MRSN).

Natalis inconspicua : Lectotype (designated here): Natalis inconspicua, Blackb. ; Australia, Blackburn Coll., B.M. 1910-236; P. 4ine (TYPE) (BMNH).

Comment on the type specimens: Notoxus porcatus: As Fabricius (1787) made no reference to specimens in his description it cannot be assumed that the nominal species-group name, Notoxus porcatus , was based on a single specimen. The 'type' specimen located in the Natural History Museum, London (BMNH) must therefore be considered a syntype (ICZN 1999: Article 73.2) and available for lectotype designation (ICZN 1999: Article 74.1).

Natalis cribricollis : At the end of his description Spinola (1844: 203) writes "Une femelle, de la collection Dupont. Mâle, inconnu" (i.e. one female/male unknown), implying that the nominal species-group name, Natalis cribricollis , was based on a single specimen. The 'type' specimen lodged at the Museo Regionale di Scienze Naturali, Turino (MRSN) is therefore considered the holotype, fixed by monotypy (ICZN 1999: Article 73.1.2). Ekis (1975), correctly, recognised this specimen as a holotype.

Natalis inconspicua: As Blackburn (1890) made no reference to specimens in his description it cannot be assumed that the nominal species-group name, Natalis inconspicua , was based on a single specimen. The 'type' specimen located in the Natural History Museum, London (BMNH) must therefore be considered a syntype (ICZN 1999: Article 73.2) and available for lectotype designation (ICZN 1999: Article 74.1).

Others: Natalis porcata Fab , Tasmania; 25-035861, 3 ex. (SAMA). Hobart, Tas., Lea; A.H. Elston Collection; K 324384 (AM). Museum Paris, Tasmanie, J. Verreaux 1844; Natalis porcata F., nec. Spin., Schenkling vid, 1901 (MNHN). Museum Paris, Tasmanie, Verreaux 3–47; Natalis porcata F., nec. Spin., Schenkling vid, 1901 (MNHN). Hobart, Tas., 28.2.20 (NMV). Terra Van Diemen, Notoxus porcatus F. (MNHN). Hobart, Tasmania, 1913-117 (BMNH). G.H. Hardy, Hobart, 17.8.13 /15 (QM). Hobart, Tas., I.1903, J.J. Walker (HEC). Launceston, Tasmania (QM). Launceston, T., F.M. Littler, 18_1_16, 2 ex. (QM). Launceston, Tasmania, 2 ex. (QM). Launceston, Tasmania, J.J. Walker, 2 ex. (BMNH). Emu Bay (MNHN). Lake Broadwater, At light, Via Dalby, SE. Qld., site 6, 22–25 Feb 1986, G. Monteith & G. Thompson (QM). Flew to outside light, Warm Stn, Newstead, Laun. TAS, Feb. 80 (ANIC). Flew to outside light, 11 pm overcast, humid, still, Newstead, Laun. Tas., 29-12-79 (ANIC). Australia, Tas, Apsley Gorge, Douglas Apsley National Park, 41°52'S 148°10'E; 6 December 1993, D.S. Horning, Jr., Under bark, Eucalyptus sp., TAS-126, 2 ex. (ANIC). Canberra, F.O.T., 1.2.1949, G.F. Hill (ANIC). Dorrigo, N.S.W., C. Oke (NMV). Victoria; Natalis cribricollis Du (MNHN). Emu Bay, Tasmania (MNHN). Australia, Tas., 2km NW Derwent Br., 730m, 24–28 Jan. 1980, wet sclerophyll, A. Newton, M. Thayer; pyrethrum spray, tree bark, at night (ANIC). Moe, Vict., 16-9-1945, 4-9-1945, 10-6-1946, C.G.L. Gooding (ANIC). Bendoc, V., Jan. 1938, F.E. Wilson, F.E. Wilson Collection (NMV). 4km E of Guyra, NSW, 4.x.1971, S. Misko (ANIC). 8km NW of Walcha, NSW, (30.59S, 151.36E), on Rd to Wollum-Walcha, 3.x.71, S. Misko, 2 ex. (ANIC). Outside light, Beechford, TAS., 19/12/1983, S. Fearn (ANIC). Hopetoun, W.A., 6 Sep.–25 Oct. 1984, A. Farman (ANIC). Ridgeway, Tas., 10/48, Co. (ANIC). Ridgeway, Tas., C. Oke, Oct. 1948 (NMV). 43.22S 146.08E, TAS, Clayton, Bathurst Harbor, 15 Jan.–20 Feb. 1991, A. Calder, W. Dressler, malaise '3, closed forest, ANIC Database No. 25 0 29657 (ANIC). 43.25S 146.10E, TAS, Melaleuca, near Bathurst Harbour, 20 Feb.–15 Mar. 1991, M. Horak, P. McQuillan, malaise #1, closed forest (ANIC). 43.25S 146.10E, TAS, Melaleuca, Bathurst Harbour, 25–29 Nov. 1991, E. Nielsen, G. Clarke, at light (ANIC). Gawler (NMV). Mt. Buffalo; Australia, C.E. Clarke Collection, B.M. 1957-24, 2 ex. (BMNH). Mt. Buffalo, V., Blackburn; 25-035815 + 25-035826 (SAMA). Mt. Wellington, S. Tasmania, 15 Jan.–6 Feb. 1913; 1,300–2,300 ft, R.E. Turner, 1913-212 (BMNH). Victoria, Hauschild; Coll. S. Schenkling; porcata Fab. ; Blackburn, Oct. 1904; Col-02928 (SDEI). Blue Mts., N.S. Wales, G.E. Bryant, 3.1.0 9 (BMNH). Blue Mts., N.S. Wales, G.E. Bryant, 1.1904 (BMNH). Tas., 4 km ESE of Richmond, 27 Mar. 1977, A.A. Calder (NMV). Harrietville, Jan 1920 (NMV). Kelso (QM). Australia-Vic., Baxter 29.9.57, D.R. Holmes lgt.

(JKCB). Australia, ne. Tasm., Mt. Barrow, I. II.1972, 40km e Launceston, leg. M. Baehr, 3 ex. (RGCM). Australia, NSW, Aspley Gorge NP, 1.– 2.12.1990, leg. M. Baehr (RGCM). Victorian Alps, Blackburn; 25-035954 (SAMA). Armidale, N.S.W., 10-V-1984, M. Coombs (QM). Rockhampton, Heyne Broth.; Col-02927 (SDEI). Australia-Vic., Red Hill, D.R. Holmes lgt. + 20.2.60, 2 ex. (JKCB). Australia, e. Tasmania, Freycinet Pen., 8.11.1972, leg. Baehr (RGCM). Murrumbidgee R., N.S. Wales, H.J. Carter; K 324381 (AM). Australia '11, TAS67, Peggs Beach Cns. Area, 8km e. Stanby, 18m, 40°50'59"S, 145°19'44"E, 25m, 3.2.2011, M. Baehr (RGCM). Ulverstone, T., Xmas 1924, R. Blackwood; K 324363 (AM). Lithgow, N.S.W., June 1924, H.E.P. Bracey; K 324357 (AM). Huon R., Tas., Lea (MTD). Franklin, Huon River, Tasm., J.J.W. (BMNH). Australia, N. Queensland. Mackay, D.S. North (BMNH). Parattah, Tas: Lea (QM). Liffey Valley, Tas., under bark on eucalyptus , 5.10.80, S. Fearn (ANIC). Under bark on standing dead eucalypt, Liffey Valley, Tas., 6/9/1981, S. Fearn (ANIC). Victoria, Barton, 2 ex. (MNHN). Rokeby. V., Nov-1965, C.G.L. Gooding (ANIC). Australia, NSW, Hanging Rock, 31°29'S, 151°12'E, 8 June 1992; D.S. Horning, Jr., Under bark Eucalyptus sp., NSW-8 (ANIC). 3km W Monga, NSW, 25 Nov. 1995, ex bark & soil at base of trees, G.W. Ulrich coll. (ANIC). Picadilly Circ., Brindabella Rng., ACT, 22 Nov. 1982, under Eucalyptus bark, J.&N. Lawrence (ANIC). Tasmania, Brighton; K 324385 (AM). Australia, SA, Pt. Lincoln NP, September Beach, 20.1.2004, M. Baehr (RGCM). Kangaroo Is, J.G.O. Tepper; K 324390 (AM) + 25-035972 (SAMA). W. Australia, Zanthus, 5.11.2001, L. Hovorka leg. (JRCP). W. Australia, Coolgardie leg., 18.X.2001, Hovorka (JRCP).

Diagnosis: The long, slender habitus with subparallel elytra is very similar to E. planipennis , but E. porcata differs from E. planipennis by the dense and large pronotal punctation; the pronotal sides are only slightly wrinkled or with coarse punctation. The tegmen is broad and not constricted in the middle, the phallic tip is round, then constricted

. Length: (247 specimens measured) Total length: 11–19 mm.

Head: Dark brown to black, with dense pilosity; with dense small and large punctation; epistomal suture more or less distinct; interocular space about 1.5 one eye width.

Thorax: Pronotal length to width ratio 1.15:1; pronotum light brown to dark brown or black; with isolated pilosity; conspicuously longer than wide; with dense, large punctation; sides slightly wrinkled or with coarse punctation; grooves of sub-basal collar less distinct, separated by a bar. Scutellum light brown to dark brown, with isolated pilosity.

Elytra: Elytral length to width ratio 3.10:1; elytra light to dark brown, with isolated pilosity; elytral base slightly wider than pronotum, subparallel; diameter of punctations in basal half slightly smaller than interstices, punctation decreasing behind middle; punctations of tenth stria distinct almost towards apex; punctations of ninth stria distinct from middle towards apex; elytral punctation with three nodules; interstices 4, 5 and 6 slightly ridged in apical half; mostly without microsculpture, partially with very weak microsculpture.

Lower surface: Dark brown to black, with dense pilosity; metaventrite and abdominal ventrites with dense punctation; procoxal cavities opened to closed posteriorly.

Legs: Light brown to dark brown, with dense pilosity and punctation, with fine wrinkles; mesotibiae bent.

Abdomen: Tegmen ventrally closed, sides open; phallic tip round, then constricted; spicular fork fused to onefourth.

Distribution: Tasmania and mainly in coastal regions of southern Australia, with a single location in Western Australia.

Seasonal occurrence: Collected during the whole year.

Figures

FIGURE 39. Eunatalis porcata: (A) Phallus lateral, (B) Phallus ventral, (C) Tegmen ventral, (D) Spicular fork, (E) Pygidium,

FIGURE 40. Distribution of Eunatalis porcata.

FIGURE 57. (A) Eunatalis pernodulosa n. sp., (B) Eunatalis planipennis, (C) Eunatalis porcata, (D) Eunatalis punctata n. sp., (E) Eunatalis schopfi n. sp., (F) Eunatalis semicostatata.