Nedyopus latus, Chen & Golovatch & Chang, 2006

Chen, Chao-Chun, Golovatch, Sergei I. & Chang, Hseuh-Wen, 2006, The millipede tribe Nedyopodini, with special reference to the fauna of Taiwan (Diplopoda: Polydesmida: Paradoxosomatidae), Journal of Natural History 39 (47), pp. 3997-4030 : 4011-4013

publication ID	https://doi.org/ 10.1080/00222930600556112
persistent identifier	https://treatment.plazi.org/id/03CC8786-FFE7-FFE9-EEB4-FBE1FD62FA5C
treatment provided by	Felipe
scientific name	Nedyopus latus
status	sp. nov.

Treatment

Nedyopus latus View in CoL , new species

( Figures 32–41 View Figures 32–41 , 81–88 View Figures 81–88 , 115–120 View Figures 115–120 )

Holotype: „ ( NCHUL), Taiwan ( R.O.C.), NanTou County, LuGu, SiiTou , bamboo forest, ca 1160–1398 m a.s.l., 31 October 1997, leg. S.-H. Wu. Paratypes: 2 „, 9♀ ( NCHUL), same locality and date, together with holotype. 4 „, 4♀ ( NCHUL), same county, RenAi, HueiSun, decayed wood in natural forest, 1664 m a.s.l., 27 December 1997, same collector. 2 „, 1♀ (NTNUL-My 46–48), Taichung County, HePing, GuGuan, JiiaBaoTai, ca 1000–1406 m a.s.l., 25 January 1988, leg. S.-H. Chen. 1♀ (JDLee20021114010), same county, LuGu, SiiTou, TuDiGongLun walk path, ca 1160– 1398 m a.s.l., 15 November 2002, leg. J.-D. Lee.

Diagnosis

Differs from congeners by the larger size, combined with the rather well-developed paraterga; an axial line is evident from the collum to the tip of the epiproct, and the gonopod femorite is rather narrow basally (see below and key).

Description Length ca 26–29 („, n 55) or 29–31 mm (♀, n 55); width of midbody metazona 3.5–4.0 („)

or 4.0–4.8 mm (♀). Holotype ca 27 mm long and 3.5 mm wide.

Coloration in alcohol dark brown to grey-brown in both sexes, prozona light grey-brown to grey-brown, with posterior parts near stricture darker grey-brown; sometimes axial line light brown to blackish; epiproct light grey-brown to light brown; paraterga light greybrown to light brown, pleurotergal region lighter grey-brown to brown, pleurosternal region lighter grey-brown to light brown, sterna light grey-brown, basal segments of legs lighter grey-brown, distal podomeres dark brown to brown.

Main somatic and gonopod characters as in N. caudatus , but width of head5segment 354,collum,2Ḳ658511514,55759510512– 15 in „, or head,collum5354,2,556,7– 15 in ♀. Antennae long, slender, reaching stricture of metatergum 4 dorsally in „ ( Figures 32 View Figures 32–41 , 81, 82 View Figures 81–88 ), a little shorter in ♀, reaching anterior part of metatergum 3. Paraterga rather well developed ( Figures 32–34 View Figures 32–41 , 81–86 View Figures 81–88 ), slightly declined, narrowed distad; calluses with four minute pits (insertion points of abraded setae) at lateral margin ( Figures 32–34 View Figures 32–41 ) on collum and pore-bearing segments, three pits on segments 2, 3, and poreless segments, sometimes poorly expressed on segment 19; calluses delimited by a sulcus both dorsally and ventrally, but a line in place of ventral sulcus on poreless segments; calluses mostly like high ridges (poreless segments) or flat but evident discs (pore-bearing segments), never spiniform caudally. Axial line evident on collum to end of segment 19 or even up to end of epiproct. Transverse sulcus reaching base of paraterga, visible on segment 19, beaded at bottom on segments 5–17, especially poorly beaded on segment 18 ( Figures 33, 34 View Figures 32–41 , 83, 84 View Figures 81–88 ). Surface sometimes transversely rugulose on metaterga ( Figure 34 View Figures 32–41 ), longitudinally rugulose at posterior end of metaterga and on metazona below paraterga, where also evidently and densely granular ( Figures 32, 33 View Figures 32–41 , 82 View Figures 81–88 ). Stricture between pro- and metaterga evidently but not very strongly beaded, general appearance of body non-moniliform; stricture slightly wider and deeper in „. Tergal setae short, pattern 2+2 anteriorly on collum, 2+2 anteriorly on segment 2, 1+1 on segment 3 (often the setae abraded), onward always abraded. Ozopores lateral, lying on callus ca one-third metatergal length from caudal edge ( Figures 33, 34 View Figures 32–41 , 83, 84 View Figures 81–88 ). Pleurosternal carinae present, like wide rounded ridges on pregonopodial segments („), with a small caudal tooth on segment 5, with a larger one on segment 6 and a particularly prominent one on segment 7, all granular, sharp and triangular, nearly reaching caudal metazonal margin („), or like narrow ridges on segments 2–4 (♀), onward increasingly poorly developed, like low bosses traceable until segment 15 or 16 in „ ( Figures 32, 33 View Figures 32–41 , 82, 84 View Figures 81–88 ) or until segment 9 in ♀, thereafter virtually missing. Epiproct long in lateral view, ratio of epiproct length to pre-epiproct length of telson 1:2.6 in „ ( Figure 86 View Figures 81–88 ), in dorsal view subtruncate, slightly emarginate; pre-apical papillae evident, rather close to apex ( Figures 35 View Figures 32–41 , 85 View Figures 81–88 ). Hypoproct straightly to roundly subtrapeziform ( Figures 36 View Figures 32–41 , 87 View Figures 81–88 ), 1+1 setae at caudal corners situated on very small knobs, sides concave.

Sterna sparsely setose, more sparsely in ♀, not modified except for a deeply notched, setose, rather high lamina between „ coxae 4 ( Figures 37 View Figures 32–41 , 88 View Figures 81–88 ), as well as paramedian bunches of strong setae between „ coxae 5–7.

Male legs 1 to those of segment 14 or 15 with tarsal brushes ( Figure 38 View Figures 32–41 ), setation gradually thinning out toward telson, becoming relatively modest after segments 15 or 16. Legs relatively long, ca 1.5 („) ( Figures 33 View Figures 32–41 , 84 View Figures 81–88 ) or 1.2 times (♀) longer than midbody height.

Gonopods ( Figures 39–41 View Figures 32–41 , 115–120 View Figures 115–120 ) complex, much as in N. caudatus , but femorite narrower at base, and distal part of l not denticulate.

Etymology

To emphasize the relatively wide paraterga.