Nomada binotata (Robertson 1903)

Gibbs, Jason, Ascher, John S., Rightmyer, Molly G. & Isaacs, Rufus, 2017, The bees of Michigan (Hymenoptera: Apoidea: Anthophila), with notes on distribution, taxonomy, pollination, and natural history, Zootaxa 4352 (1), pp. 1-160 : 56-57

Nomada binotata (Robertson 1903) (ruficornis group: bidentate mandible) (New status, new state record)

Gnathias ovatus form binotatus Robertson 1903: 175 [female lectotype, INHS]

County records: Allegan, Dickinson, Lake, Livingston, Macomb, Midland, Montcalm, Montmorency, Newaygo, Oakland, Ottawa, Wexford.

Notes. This species was originally described as Gnathias ovatus form binotatus. The name has been infrequently cited as a synonym of the nominal species ( Alexander & Schwarz 1994; Hurd 1979; Mitchell 1962; Rodeck 1951). After examining lectotypes at INHS for this and the nominal species, JG considers that N. binotata deserves to be treated as distinct from N. ovata . Nomada binotata has a distinctive dense patch of 12 or more modified setae on the metatibial apex, but N. ovata has only four. A morphospecies recorded by Goldstein & Ascher (2016) from Massachusetts as “ Nomada [Gnathias] sp. ["multispine"]” may prove to correspond with N. binotata , in which case it is likely widespread across northeastern North America. To clarify the status of this and other morphospecies in this species group, i.e. Gnathias sensu Mitchell (1962), revisionary study is required, to clarify, e.g., the status of the four “forms” of N. ovata described by Robertson (1903) [as forms of Gnathias ovatus with masculine specific epithets], octomaculata , plena, sexmaculata, and unicolor, in addition to the two reinstated here.

Material examined. Allegan Co.: (no locality), 18 May 1959, R. & K. Dreisbach (1 ♀ MSUC) ; Dickinson Co.: Channing, T 43N R30W sec. 18, 10 Jun. 1983, Malaise trap, M. Arduser (1 ♀ MSUC): Channing, T 43N R30W sec. 18, 12 Jun. 1983, Malaise trap, M. Arduser (1 ♀ MSUC): Lake Co. : (no locality), 6 May 1945, R.R. Dreisbach (1 ♂ MSUC) ; Big Star Lake , 21–27 May 1975, Malaise trap, T.A. Bowling (1 ♀ MSUC) ; Big Star Lake , 8–10 Jun 1975, Malaise trap, T.A. Bowling (1 ♀ MSUC) ; Big Star Lake , 21–24 Jun 1974, Malaise trap, T.A. Bowling (1 ♀ MSUC) ; Big Star Lake , 29 Jun-1 Jul. 1974, Malaise trap, T.A. Bowling (1 ♀ MSUC) ; Branch , 28 May-4 Jun. 1975, Malaise trap, T.A. Bowling (1 ♀ MSUC) ; Branch , 15–17 Jun. 1975, Malaise trap, T.A. Bowling (1 ♀ MSUC) ; Livingston Co.: (no locality), 6 Jun. 1943, R.R. Dreisbach (1 ♀ MSUC) ; Macomb Co.: 9 mile Rd. & Woodard Ave., 10 May 1914, W.S. McAlpine (1 ♀ MSUC) ; Midland Co.: (no locality), R.R. Dreisbach , 12–21 Jun. 1961 (1 ♀ MSUC) ; Montcalm Co.: Flat River Game Area , 14 May 1955, R.W. Hodges (2 ♀ MSUC) ; Montmorency Co.: (no locality), 1–8 Jun. 1966, window pane trap, P.C. Kennedy (1 ♀ MSUC) ; (no locality), 15– 22 Jun. 1966, window pane trap, P.C. Kennedy (1 ♀ MSUC); Atlanta , 23 May 1957, R.L. Fischer (1 ♀ MSUC) ; Newaygo Co.: 5 mi. N Newaygo , 22–30 May 1975, Malaise trap, T.A. Bowling (1 ♀ MSUC) ; Ottawa Co.: 1 mi. W Harlem , 25 May 2005, J.K. Tuell et al. (1 ♀ MSUC:RI) ; Roscommon Co.: (no locality), 21 May 1950, R.R. Dreisbach (1 ♀ MSUC): Wexford Co. : Cadillac, 11 Jun. 1983, R.L. Fischer (1 ♀ MSUC) ; T24N R9W, sec. 16, 15 Jun. 1965, J.H. Shaddy (3 ♀ MSUC); T24N R9W, sec. 16, 23–30 Jun. 1965, J.H. Shaddy (3 ♀ MSUC); T24N R9W, sec. 22, 15 Jun. 1965, J.H. Shaddy (7 ♀ MSUC); T24N R9W, sec. 22, 23–30 Jun. 1965, J.H. Shaddy (7 ♀ MSUC).